Table 1. The sample sizes of neonate/juvenile (N/J) and adult (A) sheds, ranges of number of dorsal scale rows (at anterior-middle-posterior body), ventrals, and subcaudals, and geographic distribution for 53 snake species. See the text for detailed descriptions of the classification of body patterns .
Species | Body patterns | N/J | A | No. of dorsal scale rowsa | No. of ventralsa | Pairs of subcaudalsa | Occurrence in Taiwan or adjacent islandsb |
Achalinus formosanus formosanus | Not-patterned | 0 | 1 | 27(29)-27(25)-25 | 158-184 | 61-83 (single row) | Mountain areas of Taiwan, at altitudes of 1000-2000 m |
Achalinus niger | Not-patterned | 0 | 4 | 25-25-25 | 169-185 | 51-72 (single row) | Mountain areas of Taiwan, at altitudes of 1000-2000 m |
Amphiesma stolatum | Patterned | 2 | 27 | 19-19-17 | 142-165 | 41-87 | Throughout Taiwan, Matsu, and Orchid Is., up to 500 m altitude |
Boiga kraepelini | Patterned | 0 | 6 | 23(21,25)-21(19,23)-17(15) | 212-250 | 115-158 | Throughout Taiwan, up to 1500 m altitude |
Not-patterned | 0 | 4 | |||||
Bungarus multicinctus multicinctus | Patterned | 13 | 11 | 15(16,17)-15(16)-15 | 198-250 | 26-65 (single row) | Throughout Taiwan, Penghu, Kinmen, Matsu, Xiaoliuqiu, and Gueishan Is., up to 1000 m |
Not-patterned | 0 | 51 | |||||
Calamaria pavimentata pavimentata | Not-patterned | 0 | 2 | 13-13-13 | 167-192 | 13-23 | Throughout Taiwan and Orchid Is., up to 1500 m altitude |
Cyclophiops major | Patterned | 14 | 0 | 15-15-15 | 155-189 | 61-97 | Throughout Taiwan, up to 1000 m altitude |
Not-patterned | 0 | 18 | |||||
Daboia siamensis | Patterned | 4 | 57 | 29(27,31)-29(27,31,33)-21(23) | 151-169 | 40-54 | Southern and south-eastern Taiwan, up to 500 m altitude |
Deinagkistrodon acutus | Patterned | 1 | 8 | 21(22,23)-21(23)-17(18,19) | 152-176 | 41-63 | Throughout Taiwan, at altitudes of 150-1500 m |
Elaphe carinata yonaguniensis | Patterned-type I | 12 | 0 | 23(21,25)-23(21,25)-19(17) | 209-230 | 78-99 | Throughout Taiwan and adjacent islands, up to 2000 m altitude |
Patterned-type II | 1 | 7 | |||||
Elaphe taeniurus friesi | Patterned-type I | 90 | 17 | 25(23)-23(21,25)-19(17) | 243-262 | 101-122 | Throughout Taiwan, up to 2000 m altitude |
Patterned-type II | 0 | 102 | |||||
Euprepiophis mandarinus | Patterned-type I | 0 | 11 | 23(21,25)-23(21)-19(17,21) | 181-238 | 49-76 | Northern, Central and Eastern Taiwan, at altitudes of 1000-2500 m |
Patterned-type II | 0 | 3 | |||||
Gonyosoma frenatum | Patterned | 3 | 0 | 19(21)-19(17)-15(13) | 200-227 | 108-151 | Southern and Southeastern Taiwan, up to 1500 meters altitude |
Not-patterned | 0 | 2 | |||||
Hebius miyajimae | Patterned | 0 | 2 | 19-19-17 | 141-152 | 87-92 | Mainly in northern Taiwan, at altitudes of 500-1600 m |
Hebius sauteri | Not-patterned | 1 | 4 | 17-17-17 | 120-147 | 65-92 | Throughout Taiwan, up to 1500 m altitude |
Hebius sp. (“Paiwan Keelback”) | Not-patterned | 1 | 0 | 19-19-17? | - | - | Limited areas of southern or eastern Taiwan, between 500 and 1300 meters altitude |
Hypsiscopus plumbea | Not-patterned | 3 | 8 | 19-19-17(15) | 122-136 | 23-47 | Western and northern Taiwan, including Gueishan Is., up to 500 m altitude |
Indotyphlops braminus | Not-patterned | 0 | 8 | 20-20-20 | 300-303 | 8-14 | Throughout Taiwan and adjacent islands, up to 500 m altitude |
Laticauda colubrina | Patterned | 0 | 4 | 25-23,25-21 | 225-245 | 33-44 | Orchid Is., Green Is., eastern or southern coast of Taiwan |
Laticauda laticaudata | Patterned | 0 | 8 | 19-19(21)-17 | 215-252 | 33-47 | Orchid Is., Green Is., eastern or southern coast of Taiwan |
Laticauda semifasciata | Patterned | 0 | 10 | 23-23-19(21) | 178-203 | 32-43 | Orchid Is., Green Is., eastern or southern coast of Taiwan |
Lycodon rufozonatus rufozonatus | Patterned | 5 | 12 | 17(19,21)-17(19)-15(17) | 184-225 | 45-95 | Throughout Taiwan and Matsu, up to 2000 m altitude |
Not-patterned | 0 | 34 | |||||
Lycodon ruhstrati ruhstrati | Patterned | 38 | 7 | 17(19)-17-15 | 193-233 | 64-116 | Throughout Taiwan, up to 1500 m altitude |
Not-patterned | 0 | 7 | |||||
Macropisthodon rudis rudis | Patterned | 3 | 3 | 23-23(21,25)-19 | 123-156 | 37-65 | Throughout Taiwan, at altitudes of 500-1500 m |
Not-patterned | 0 | 2 | |||||
Myrrophis chinensis | Type I | 2 | 0 | 23(25)-23-21(17,19) | 131-155 | 35-52 | Limited areas of Taipei, Taoyuan, Nantou, Kenting, and Kinmen, at altitudes up to 500 m |
Type II | 0 | 9 | |||||
Naja atra | Type I | 54 | 0 | 23(21,25,27)-21(19,20)-15(14,13) | 158-185 | 38-53 | Throughout Taiwan and Matsu, up to 1000 m altitude |
Type II | 0 | 138 | |||||
Oligodon chinensis | Patterned-type I | 2 | 0 | 17-17-15 | 158-206 | 40-73 | Matsu |
Patterned-type II | 0 | 2 | |||||
Oligodon formosanus | Patterned-type I | 7 | 0 | 19-19(17)-17(15) | 154-189 | 39-60 | Throughout Taiwan and adjacent islands, up to 1000 m altitude |
Patterned-type II | 2 | 19 | |||||
Oligodon ornatus | Patterned | 0 | 3 | 15-15-15 | 156-182 | 27-44 | Northern and Central Taiwan, at altitudes of 500-1500 m |
Oreocryptophis porphyraceus kawakamii | Patterned-type I | 2 | 0 | 19(18,17)-19(17)-17(15) | 194-214 | 47-75 | Throughout Taiwan, up to 2200 m altitude |
Patterned-type II | 0 | 7 | |||||
Ovophis makazayazaya | Patterned | 0 | 7 | 25~31-25~29- 21 | 144-155 | 39-54 | Northern and western Taiwan, at altitudes of 500-2200 m |
Not-patterned | 0 | 3 | |||||
Pareas atayal | Not-patterned | 0 | 11 | 15-15-15 | 174-188 | 71-79 | Northern Taiwan, up to 1500 m altitudes |
Pareas formosensis | Not-patterned | 0 | 10 | 15-15-15 | 170-180 | 69-82 | Throughout Taiwan except for the north-eastern tip of the island, up to 2000 m altitudes |
Pareas komaii | Not-patterned | 0 | 7 | 15-15-15 | 162-182 | 60-76 | Central, southern, and eastern Taiwan, up to 2000 m altitudes |
Plagiopholis styani | Patterned | 0 | 5 | 15-15-15 | 102-126 | 21-32 | Very limited areas around Taipei and the Northern Cross-Island Highway, at altitudes of 500-1100 m |
Protobothrops mucrosquamatus | Patterned | 3 | 37 | 25~29-25(21,27,29)-17~21 | 194-233 | 70-108 | Throughout Taiwan, up to 1500 m altitude |
Psammodynastes pulverulentus papenfussi | Patterned | 0 | 11 | 17-17-15(13) | 161-177 | 58-79 | Throughout Taiwan and Orchid Is., up to 1500 m altitude |
Pseudoxenodon stejnegeri stejnegeri | Patterned | 0 | 2 | 19(17,21)-17-15(14) | 150-162 | 42-66 | Throughout Taiwan, at altitudes of 800-2500 m |
Not-patterned | 0 | 3 | |||||
Ptyas dhumnades | Patterned-type I | 2 | 0 | 16-16(14)-14 | 186-217 | 91-188 | Throughout Taiwan, up to 2000 m altitude |
Patterned-type II | 0 | 3 | |||||
Ptyas korros | Patterned | 2 | 0 | 15-15(13,14)-11 | 156-184 | 100-154 | Throughout Taiwan, up to 1000 m altitude |
Not-patterned | 0 | 2 | |||||
Ptyas mucosa | Patterned-type I | 25 | 0 | 19(17,21)-17(16)-14(13,15) | 170-213 | 94-143 | Throughout Taiwan, Kinmen, and Matsu, up to 1500 m altitude |
Patterned-type II | 0 | 43 | |||||
Python bivittatus bivittatus | Patterned | 1 | 18 | 53~64-64~72-40~44 | 255-263 | 63-71 | Kinmen |
Rhabdophis formosanus | Patterned | 1 | 15 | 19-19-17(15) | 161-171 | 74-89 | Mountain areas of Taiwan, at altitudes of 1500-3000 m |
Rhabdophis swinhonis | Patterned | 1 | 20 | 15(17)-15-15 | 124-165 | 49-74 | Throughout Taiwan, up to 1800 m altitude |
Sibynophis chinensis chinensis | Patterned | 0 | 4 | 17-17-17 | 164-187 | 171-208 | Throughout Taiwan and Gueishan Is., up to 1500 m altitude |
Sinomicrurus hatori | Patterned | 0 | 3 | 13-13-13 (occasionally 15) | 223-245 | 30-34 | Northern and eastern Taiwan, up to 2200 m altitude |
Sinomicrurus macclellandi swinhoei | Patterned | 0 | 1 | 13-13-13 | 207-240 | 32-41 | Northern and western Taiwan, at altitudes up to 1000 m |
Sinomicrurus sauteri | Patterned | 0 | 2 | 13-13-13 (occasionally 15) | 234-268 | 27-36 | South and central Taiwan, up to 2500 m altitude |
Sinonatrix annularis | Patterned | 10 | 0 | 19-19-17 | 136-167 | 47-74 | Very limited areas of Taipei, and Taoyuan, at altitudes up to 1000 m |
Not-patterned | 0 | 9 | |||||
Sinonatrix percarinata | Patterned | 4 | 3 | 19-19-17 | 131-160 | 44-87 | Throughout Taiwan, at altitudes up to 1500 m |
Not-patterned | 0 | 2 | |||||
Trimeresurus gracilis | Patterned | 7 | 33 | 25~27-21~19-15~17 | 144-149 | 43-53 | Central mountains in Taiwan, at elevations of 2000-3500 m |
Not-patterned | 0 | 5 | |||||
Trimeresurus stejnegeri | Not-patterned | 4 | 52 | 21(22~25)-21(23)-15 | 154-178 | 43-80 | Throughout Taiwan, Green Is. and Orchid Is., up to 2000 m altitude |
Xenochrophis piscator | Not-patterned | 2 | 9 | 19-19(17)-17(15) | 121-152 | 42-90 | Throughout Taiwan and Kinmen, up to 500 m altitude |
aThe numbers of dorsal scale rows, ventrals, and subcaudals are mainly summarized from previous reports (see the text). The data of Hebius sp. are measured from a uid-preserved specimen (voucher number: NIU-HE-001). (Maki 1931; Pope 1935; Wang and Wang 1956; Kuntz 1963; Ota and Toyama 1989; Tu et al. 1990; Zhao et al. 1998; Ota et al. 1999; Zhao 2006; You et al. 2013, 2015). bThe geographical distributions of snakes in Taiwan and adjacent islands are mainly according to Shang et al. (2009).