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. 2019 Mar;22(3):225–237. doi: 10.22038/ijbms.2019.32806.7897

Pharmacological activities of gallic acid and its derivatives in different diseases

Disease category Compound name Model Effects References
Anti-inflammatory
Gallic acid In vitro: LPS-induced inflammation in A549 lung cancer cells
In vivo: LPS-induced inflammation in mice
In vitro: ↓HAT, ↓p300 & CBP acetyltransferase, p300-mediated RelA acetylation, NF-κB–regulated antiapoptotic & cell survival genes expression, p300-induced p65 acetylation, LPS-induced p65 translocation to the nucleus, ↑cytosolic IĸBα
In vivo: ↓p65 acetylation, IFN-,
IL-6, IL-1β & NF-κB–regulated antiapoptotic genes expression
(72)
Gallic acid In vitro: zymosan-induced damage in human PMN
In vivo: zymosan-induced acute food pad swelling in mice
In vitro: interference with PMN function, ↓MPO & reduction rate of cytochrome c
In vivo: ↓footpad swelling size
(71)
Gallic acid ethyl ester Acetic acid-induced abdominal constriction, formalin-induced nociception, capsaicin-induced nociception, rat paw hyperalgesia induced by substance-P, bradykinin, PGE2 or carrageenan ↓Acetic acid-induced abdominal constrictions, ↓formalin-induced licking,
↓hyperalgesia induced by substance P & bradykininNo significant change in capsaicin-induced nociception
(70)
Bergenin (C-glycoside of 4-O-methyl gallic acid) Mycobacterium tuberculosis-induced inflammatory arthritis in mice ↓Inflammatory arthritis, ↓IL-2, IFN-γ, TNF-α, IL-4 & IL-5 (69)
Gallic acid In vitro: AEGs- treated rabbit chondrocytes
In vivo: collagenase-induced knee osteoarthritis in rabbit
In vitro: ↓ROS, collagen II & aggrecan degradation, NO, i-NOS, COX-2, PGE2
↑GSH, SOD
In vivo: ↓knee Mankin’s score
(73)
Cardiovascular
Gallic acid ISO-induced myocardial infarction in rats ↓Myocardial injury, ↓TC, TG, LDL-C, VLDL-C, MDA, ↑HDL-C, CAT & GPx, ↑membrane-bound Na+/K+, Ca2+ & Mg2+ ATPase (85)
Gallic acid ISO-induced cardiotoxicity in rats ↓CK-MB & LDH,
↓lysosomal membrane damage, LPO, ↑GSH
(86)
Gallic acid Lindane-induced cardiotoxicity ↓CK, LDH & LPO, ↑GSH, SOD, GPx & GST, ↑membrane bound Na+/K+ & Mg2+ ATPase & ↓ Ca2+ATPase (87)
Gallic acid Evaluation of antioxidant enzymes in the heart of male Sprague-Dawley rats ↑Cardiac SOD, GPx, CuZnSOD, CAT, GSH/GSSG ratio, heme oxygenase-1 & Nrf2 (88)
Gallic acid AGEs-induced cardiac remodeling in rats ↓Cardiac fibrosis, ↓TNF-α, TGF-β, MMP-2 & MMP-9 (89)
Gallic acid STZ-induced myocardial dysfunction in diabetic rats ↓CK-MB, ↓LDH, LPO, LDL-C & VLDL-C, ↓MBP, SBP & bradycardia, collagen content,
↑CAT, SOD, GSH
(90)
Gallic acid Isoproterenol-induced myocardial infarction in rats ↓CK-MB, ↑SOD, CAT, GPx, GST, GSH, Vit C & E, ↓troponin-T, LDH-1 & LDH-2 (58)
Gallic acid Fructose-enriched-diet-induced cardiac fibrosis ↓BP, HOMA-IR, ↓NADPH oxidase subunits gp91 phox & p22 phox, ↓collagen I & osteopontin (91)
Gallic acid Al2O3-induced myocardial injury ↓LDH, CPK, CK-MB, TG, LDL, TNF-α & MDA, ↑HDL, GSH, SOD & CAT (92)
Gallic acid Alloxan-induced diabetes & endothelial dysfunction ↓MDA, ↑TAC & histamine vasodilatory response of mesenteric vascular bed (93)
Gallic acid L-NAME-induced hypertension ↓SBP, LV wall thickness & cardiac fibrosis, ↓hypertrophy markers, ↓HDAC1 & 2 (94)
Gallic acid Cyclophosphamide-induced cardiorenal dysfunction ↓MDA & H2O2, ↑CAT, GST, GSH & GPx (95)
Gastrointestinal
Gallic acid CCl4-induced hepatotoxicity in Charles Foster rats & Swiss albino mice ↓Sleep time & paralysis time, ↓LPO, ↑hepatic amidopyrine-N-Demethylase, aniline & membrane-bound hepatic glucose-6-phosphatase activity, ↓hepatic TAG (53)
Gallic acid Hepatic ischemia
& reperfusion injury in rats
↓ALT, AST & LDH activities, ↑CAT & GPx, ↓MDA (57)
n-propyl gallate Isolated perfused rat liver ↓Gluconeogenesis, pyruvate carboxylation,
glucose output inhibition
(96)
Gallic acid Brush border disaccharidases inhibition in rats, LACA/L mice, BALB/c mice & rabbit ↓Sucrase, maltase, trehalase & lactase activity (97)
Gallic acid Primary HSC & hepatocytes Cytotoxicity to HCS but not hepatocytes,
↑intracellular Ca2+ & calpain activity
(98)
Gallic acid Ethanol-induced pancreatic injury in rats ↑Cathepsin B activities, ↓cathepsin B & L enzymes release, cytosolic/lysosomal ratio of cathepsin B & L, pancreatic tissue injury (99)
Gallic acid Ethanol-induced liver damage in rats ↓AST, ALT, LDH activity, ↑paraoxonase & arylesterase activity (54)
Gallic acid Gastric mucosal lesions caused by ischemia-reperfusion injury in rats ↓Total area of gastric lesions, ↓caspase-3 & i-NOS (47)
Gallic acid In vitro: rat gastric epithelial cells
In vivo: indomethacin & diclofenac-induced gastropathy
In vitro: ↓mucosal cell death, ↑mitochondrial dehydrogenases
In vivo: ↓mitochondrial protein carbonyl formation, ↓LPO & caspase-9 activation, ↑thiol content & MMP
(48)
Tryptamine-gallic acid In vitro: rat gastric epithelial cells
In vivo: indomethacin-induced gastropathy
In vitro: ↓intramitochondrial ROS generation

In vivo: ↓mucosal cell death, gastropathy,
mitochondrial protein carbonyl formation,
↓LPO, bcl-2 expression & caspase-9 activation, ↑thiol content & bax expression
(100)
Gallic acid DSS-induced experimental colitis in mice ↓DAI & colon shortening, ↓IL-21, IL-23, MDA, ↑SOD, GPx, CAT, GR, Nrf2, UDP-GT & NQO1 (50)
Gallic acid DSS-induced colitis in mice ↓MPO activity, i-NOS, COX-2, p65-NF-κB & IL-6/p-STAT3Y705 activation (51)
Gallic acid Paracetamol-induced liver damage in mice ↓ALT, AST, ALP, & ↓TNF-α, ↑SOD, CAT, GSH, GPx & GST (101)
Trimethylgallic acid esters CCl4-induced liver damage in rats ↓Vacuole formation, inflammation & necrosis, ↓AST, ALT, TG, TC, LPO & ↓TNF-α, ↑SOD, CAT & GSH (102)
Gallic Acid Aspirin + pyrolus ligation-induced gastric ulcer in rats ↓Ulcer index, gastric juice volume, free & total acidity, total protein, carbohydrates concentration, ↑SOD, CAT, GSH, GPx, GR & glucose-6-phosphate dehydrogenase (103)
Gallic acid Bromobenzene-induced liver injury in rat ↓Aniline hydroxylase & AMND activity,
↓LPO, ↑epoxide hydrolase activity
(52)
Gallic acid CCl4-induced liver fibrosis in mice ↓Liver fibrosis, HA, MDA, ALT, AST & GGT (104)
Gallic acid & piperine Beryllium-induced hepatorenal dysfunction in rats ↓Bilirubin, Cr, LDH, GGT, LPO, AST, ALT, ALP, ↑GSH, SOD & CAT (105)
Gallic acid Lead-induced toxicity in blood, liver & kidney of rats ↓LPO & carbonyl, prevention of body weight loss, ↑ALA-D activity, ↑SOD, CAT & GSH (56)
Gallic acid & ellagic acid LPS-induced liver injury ↓ALT, AST & i-NOS expression (106)
Gallic acid CCl4-induced chronic liver injury in rats ↓ALT, AST & MDA, ↑SOD, CAT, GSH, GR, GPx & GSH/GSST (107)
Gallic acid Lindane-induced hepatorenal toxicity in rats ↓ALT, AST, ALP, LPO, creatinine & urea, ↑GSH, CAT, SOD, GPx & GST (108)
Gallic acid Beryllium-induced hepatorenal toxicity ↓AST, ALT, ALP, LPO, AMND, ↑GSH, CAT, SOD, GPx & GST, ↓Cr & urea (109)
Gallic acid Cyclophosphamide-induced hepatotoxicity in rats ↓AST, ALT, MDA, ↑GSH, CAT, SOD & GST (55)
Gallic acid Indomethacin-induced gastric ulcer in Swiss albino mice ↑Ulcer healing, ↓PGE2 synthesis, ↑e-NOS/i-NOS ratio (49)
Metabolic Gallic acid Diet-induced obesity in mice ↓TAG & FBS, ↓adipocyte size in the epididymal white adipose tissue, ↑PPAR expression, ↑Akt signaling pathway activity, ↓glucose tolerance & lipid metabolism (59)
Gallic acid High-fat-diet- & -STZ-induced type 2 diabetes in rats ↓Body weight gain, FBS & FPI, ↑adipose tissue insulin sensitivity, Cytoprotective action on pancreatic β-cell, ↑PPARγ expression in treated tissue, liver & skeletal muscle, ↑insulin-dependent glucose transport, ↑interactions with the GLUT4, GLUT1, PI3K & p-Akt, ↓adipogenesis (60)
Gallic acid High-fat-diet-induced dyslipidemia, hepatosteatosis & oxidative stress in rats ↓Obesity, liver weight, peritoneal & epididymal adipose tissue weights, ↓serum TAG, phospholipid, TC, LDL-C, insulin & leptin, ↓lipid droplets size, ↓hepatic TAG & cholesterol, ↓oxidative stress & GSSG, ↑GSH, GPx, GR & GST (61)
Gallic acid High-fructose-diet-induced diabetes ↑Glucose uptake activity, ↓AUCglucose & HOMA-IR, ↓C-peptide, fructosamine & cardiovascular risk index, ↑IR, IR-1, PI3K, Akt/protein kinase B & GLUT-2, ↓F-1,6-BP, ↑hexokinase, PFK & aldolase (62)
Gallic acid STZ-induced diabetic rats ↑Vit C, ↓GSH, ↓LPO, ↑free radical scavenging property, Fe2+ chelating ability & Fe3+ reducing property, ↑CAT, GST, δ-aminolevulinic acid dehydratase & LDH, ↓purinergic enzymes (63)
Gallic acid STZ-induced diabetic Wistar rats ↓FBS, regeneration of β-cells, ↓TC, TAG, LDL-C, urea, uric acid, creatinine, ↑FPI, C-peptide & glucose tolerance restored the total protein, albumin & body weight (110)
Gallic acid Fructose-induced metabolic syndrome & cardiac fibrosis in rats ↓Insulin resistance, ROS & NADPH overproduction, collagen I & osteopontins (98)
Gallic acid In vitro: porcine pancreatic lipase kit
In vivo: high-fat-diet-induced obesity in mice
In vitro: ↓pancreatic lipase activity
In vivo: ↓weight gain, ↑feces neutral fat
(111)
Gallic acid STZ-induced diabetes in rats ↑FPI, hepatic hexokinase activity, CAT, SOD, GPx, ↓FBS, HbA1C, G6PD & fructose-1, 6-bisphosphatase, LPO (112)
Gallic acid STZ-induced diabetes in rats ↓FBS, HbA1C, LPO, ↑FPI, Vit C, SOD, CAT, GSH, GR, GST, GPx, HMG-CoA reductase activity (113)
Gallic acid Alloxan-induced diabetes in rats ↓FBS, ↑FPI, GSH, GPx, CAT, SOD & osmotic fragility of RBCs (114)
Gallic acid STZ-induced diabetes in rats ↓FBS, brain LPO, SOD, CAT, GR, GST, GPx, brain lipids (37)
Gallic acid Chromium-induced thyroid dysfunction ↓SOD & GST up-regulation, ↓NO, i-NOS, TNF-α, IL-6 & COX-2 (115)
Gallic acid In vitro: high glucose toxicity in NRK 52E rat proximal tubular epithelial cells
In vivo: high fat diet/STZ- induced diabetes in rats
In vitro: ↓p38 MAPK, NF-κB activation
In vivo: ↓FBS, HbA1C, HOMA-IR, body weight, Cr, Cr clearance, BUN, IL-1β, IL-6, TNF-α & ↓MDA, ↓renal p38 MAPK, NF-κB activation, TGF-β, fibronectin, ↑GSH, GSST, GSH/GSST ratio, GR, CAT, SOD & GPx
(116)
Gallic acid STZ-induced diabetes & oxidative stress in rats ↓ROS & lipid peroxidation, ↑SOD & δ-ALA-D, CAT, GST & vit C (117)
Neuropsychological
Gallic acid 6-Hydroxydopamine induced oxidative stress in rats ↑Passive avoidance memory, ↑TTM, GPx, ↓MDA (65)
Gallic acid STZ-induced memory deficits & oxidative stress in rats ↑Passive avoidance & spatial memory, performance, ↑TTM, SOD, GPx & CAT, ↓MDA (118)
Gallic acid EPM in rats ↑Time spent & entries in the open arms of EPM, ↓locomotor activity, involvement of 5-HT1A receptors (119)
Gallic acid Sodium fluoride-induced oxidative stress in rat brain ↓LPO, ↑SOD & GSH (120)
Gallic acid STZ-induced oxidative damage in rat brain ↓MDA, ↑TTM, CAT, SOD & GPx, ↑ Na+/K+, Ca2+ & Mg2+ ATPases activity (121)
Gallic acid Spinal cord injury-induced oxidative stress in rat ↓LPO, ROS, nitrite, NF-kB & COX-2
↑GSH, CAT, SOD & GPx
(122)
Gallic acid (as chitosan nanoparticles) Scopolamine-induced amnesia & locomotor activity ↓Transfer latency in the EPM test, ↑spatial learning & memory in MWM, ↓AChE activity, (66)
Gallic acid Tyrosine hydroxylase Gal4/UAS-X RNAi Drosophila melanogaster model of Parkinson's disease ↑Locomotor activity, protection of dopaminergic neurons, ↑life span & climbing abilities (123)
Gallic acid Cyclophosphamide-induced neurotoxicity in rats ↓Neurotoxicity, ↓cerebellar & cerebral MDA & nitrite, ↑CAT, GST & SOD (55)
Gallic acid Reserpine-induced vacuous chewing movements in rats ↓Vacuous chewing movements (124)
Gallic acid Lead-induced locomotor damage & brain oxidative stress in rats ↑Locomotor & exploratory activities by attenuating crossing & rearing time, ↓brain levels of Pb, ↑SOD & ↑GSH (125)
Gallic acid Sodium nitroprusside oxidative stress-induced mitochondrial impairment ↓NO level, ↓mitochondrial protein tyrosine nitration, ↓LPO, ↓protein carbonyl, ↑GSH & ↓MPT (126)
Gallic acid In vitro: sodium hydrosulfite-induced mitochondrial dysfunctions in SH-SY5Y cells
In vivo: cerebral ischemia/reperfusion-induced by middle cerebral artery occlusion
In vitro: protects against cytotoxicity of SH-SY5Y cells, ↓mitochondrial dysfunction, ↓level of mitochondrial ROS by ↓MitoSOX-fluorescence intensity, ↓intracellular DCF-fluorescence intensity, ↓intracellular MDA, by modulating mitochondrial dysfunctions by ↑oxygen consumption
In vivo: ↓total infarct volume
(127)
Gallic acid (as chitosan nanoparticles) FST & TST in rat ↓Immobility in FST & TST, ↓MAO-A activity & MDA, ↑GSH & CAT (128)
Gallic acid Aβ-induced toxicity in cultured rat cortical neurons ↓Apoptotic neuronal death, ↓(Ca2+)c elevation & ROS formation, ↑glutamate release (64)
Gallic acid H2O2-induced apoptosis in rat pheochromocytoma PC12 cells Gallic acid & EGCG: ↓cell viability
methyl gallate: ↑cell viability, ↓mitochondrial depolarization, caspase-9 activation & DNA degradation
(68)
Gallic acid Immobilization-induced Swiss male albino mice ↓Plasma nitrite in both unstressed & stressed mice, ↓plasma corticosterone, ↓n-NOS activity, ↓anxiety in behavioral tests (129)
Gallic acid Global ischemia/reperfusion in Wistar rats ↑Gait performance, sensorimotor disorders, & hypoalgesia (high dose), ↑passive avoidance memory (low dose), improvement in behavioral motor activity tests (130)
Gallic acid Experimental sciatic nerve crush in rats Improved motor coordination & SNCV sciatic nerve conduction velocity, ↑delayed foot lifting (131)
Gallic acid Aβ-induced AD in rats Improved LTP amplitude & area under the curve,
↑PS Amp, ↓Aβ plaque
(132)
Gallic acid H2O2-induced apoptosis in rat pheochromocytoma PC12 cells ↓Cell viability, ↑PARP cleavage, ↑JNK phosphorylation, ↓Bcl-2 (67)
Gallic acid STZ-induced cerebral oxidative stress in rats ↑Weight loss, ↓hyperglycemia, HbA1C, LPO, AChE & purinergic enzymes, ↑radical scavenging & Fe2+ chelating ability, Vit C, GSH, CAT, GST, cerebral LDH & Na+/K+-ATPase activity (133)
Gallic acid In vitro: Aβ-induced neurotoxicity in murine microglial BV-2 cells & neuroblastoma Neuro-2A cells
In vivo: Aβ-induced AD in ICR mice
In vitro: ↓RelA acetylation & cytokine production, cell death, ↑viability of Neuro-2A, ↓memory deficits in Ab peptide-induced mice
In vivo: ↓cytokine production, neuronal cell death, nuclear NF-κB & IL-1β
(134)
Gallic acid Chronic cerebral hypoperfusion-induced cognitive deficit & brain oxidative damage in rats ↑Spatial memory, ↑TTM & GPx, ↓LPO (135)
Gallic acid & its derivatives 6-OHD-induced neurotoxicity in human SH-SY5Y neuroblastoma cells ↓Neurotoxicity, ↑GSH, ↓GSSG, ↓elevation in (Ca2+)I (136)
Oral health Gallic acid Streptococcus sobrinus 6715-induced enamel caries in rats ↑Remineralization of enamel caries lesions, residual first molar enamel volume & mineral density values, ↓severity of molar enamel caries (137)
Radiation-induced toxicity Gallic acid Whole body γ-radiation exposure in mice ↑Rate of DNA repair process in peripheral blood leukocytes, bone marrow cells, & splenocytes, ↑GPx, GSH, ↓mortality, weight loss & LPO (82)
Gallic acid In vitro: rat liver microsomes & plasmid pBR322 DNA exposed to γ-irradiation
In vivo: whole body γ-irradiation in mice
In vitro: ↓LPO in rat liver microsomes, ↓DNA damage in plasmid
In vivo: ↓DNA damage in leukocytes
(81)
Respiratory Gallic acid Bleomycin-induced pulmonary fibrosis in rats ↓Lesions & fibrosis, collagen content, hydroxyproline accumulation, LPO, ↓TNF-α & IL-1β, ↑GPx activity & TTM (80)
Urinary

Gallic acid Doxorubicin-induced chronic kidney disease in rats ↑Albumin, ↓AST, ↓ALT, ↓TG, ↓cholesterol, ↓LPO, ↓BUN (79)
Gallic acid Glyoxal-induced renal fibrosis in rats ↓Renal fibrosis, ↓BUN, ALP, collagen I & III, MMP-2 & -9, NOx & ROS, ↑SOD (78)
Gallic acid Ferric nitriloacetic acid-induced renal toxicity in rats ↓Renal toxicity & cell proliferation, BUN, H2O2, renal microsomal LPO & quinone reductase, ↑CAT, xanthine oxidase, GPx, GST & G6PD (77)
Gallic acid Cisplatin-induced nephrotoxicity in rats ↓LPO, ROS, Cr, urea, uric acid, arginase activity, ↑SOD, CAT, GSH & GPx (75)
Gallic acid Experimental renal ischemia-reperfusion in rats ↓BUN, Cr, MDA (74)
Urogenital Gallic acid Cyclophosphamide-induced toxicity in testis & epididymis of rats ↓Reproductive toxicity, nitrite, H2O2 & MDA ↑SOD, GST, FSH, LH & testosterone (55)
Gallic acid Cyclophosphamide-induced toxicity in testis & epididymis of rats ↓MDA, NO, H2O2, ↑GSH, GPx, SOD, CAT & testosterone (76)
Gallic acid STZ-induced oxidative stress in testis of rats ↑SOD & CAT, ↓MDA, TNF-α, VEGF & NOS2 (138)
Dermal Gallic acid In vitro: normal human dermal fibroblasts exposed to UVB
In vivo: hairless mice exposed to UVB
In vitro: ↓transcription factor activation protein 1 activity
In vivo: ↓dryness, skin thickness, wrinkle formation & MMP-1, ↑elastin, type I procollagen & TGF-β1
(84)
Gallic acid In vitro: murine melanoma B16F10 cells
In vivo: zebrafish,
UVB-induced hyperpigmentation in mice ear
In vitro: ↓melanin production & tyrosinase activity, melanogenesis regulatory genes, activation of the ERK pathway, involvement of AKT/GSK3b & PKA/CREB signaling
In vivo: ↓body pigmentation in zebrafish,
↓hyperpigmentation of ear skin, inflammation,
melanocytes activation & melanogenic genes
(83)
Malignancy Gallic acid DMH-induced colon carcinogenesis in male Wistar rats ↑SOD, GSH, GR, GPx, & CAT activity, LPO modification (39)
Gallic acid DMH-induced colon carcinogenesis ↑Activity of phase I enzymes (cyt. P450 & cyt. b5), ↓activity of phase II enzymes (GST, DTD & GGT) (139)
Isobutyl gallate-3,5-dimethyl ether (IGDE) &
methyl gallate-3,5-dimethyl ether (MGDE)
In vitro: EAT & LLC1 cells
In vivo: EAT cells /BALB/c mice & LLC1 cells /C57bl/6 mice
In vitro: no significant cytotoxic effects
In vivo: EAT cells ↑Survival (IGDE>MGDE), NK cells cytotoxicity
In vivo (LLC1): ↓tumor size (IGDE>MGDE)
(44)
Gallic acid In vitro: HL-60 human promyelocytic leukemia
In vivo: athymic nude mice model
In vitro: induction of G1 cell cycle arrest, ↓cyclin D1, CDK4, cyclin E, CDK2, & cyclin A, ↑p27KIP expression
In vivo: ↓Tumor progression
(45)
Gallic Acid Diethylnitrosamine-induced hepatocellular carcinoma in rats ↓Tumor size, AFP & CEA, ↓serum AST, ALT, ACP, ALP, LDH, GGT, ↓liver AgNORs & PCNA (46)
Gallic acid In vitro: human NCSLC NCI-H460 cells
In vivo: mouse NCI-H460 xenograft model
In vitro: ↓viability, induction of G2/M phase cell cycle arrest, ↑intracellular Ca2+, CDK1 activity, caspase-3, caspase-8 & caspase-9 activation, ↓ΔΨ
In vivo: ↓tumor size
(140)
Gallic acid In vitro: LL-2 mouse lung cancer cells
In vivo: LL-2 lung cancer cells transplanted in mice
In vitro: ↓viability
In vivo: ↓tumor size, ↑number or apoptotic cells in tumor, synergistic effects in combination with cisplatin
(141)
Gallic acid & methyl gallate two-stage skin carcinogenesis in ICR mice ↓average number of papillomas per mouse (142)
Gallic acid 7,12-DMBA/croton oil-
induced two-stage skin carcinogenesis in Swiss albino mice
↓time of appearance & average number of papillomas per mouse, tumor incidence, ↓LDH total activity& LDH-isoenzymes, LPO, MMP-2 & MMP-9 activity & expression, ↑GST, SOD, CAT activity & GSH, synergistic effect with 5-FU (40)
Gallic acid In vitro: cell-free kinases, primary HUVECs, primary human dermal LECs, human HT29 colon carcinoma cells & MT-450 rat mammary carcinoma cells
In vivo: MT-450 tumor-bearing rats
In vitro: slight inhibition of RTKs, ↓VEGF-induced autophosphorylation of VEGFR-2 in HT29 cells, ↓proliferation & ↑apoptosis in all cell lines
In vivo: ↓tumor angiogenesis, ↑metastasis
(143)
Pyrogallol In vitro: MCF10DCIS.com cells
In vivo: xenograft mouse model of MCF10DCIS.com
In vitro: induction of S phase cell cycle arrest
↑ROS
In vivo: ↓tumor size, IR, IRS1, IGF-1R, p70S6K, & ERK phosphorylation, ↓IL-1β, involvement of AMPK & AKT/mTOR signaling
(43)

i-NOS: nitric oxide synthase; IL-2: interleukin-2; IFN-γ: interferon-γ; TNF-α: tumour necrosis factor-α; IL-4: interleukin-4; IL-5: interleukin-5; IL-1β: interleukin-1β; COX-2: cyclooxygenase-2; IL-6:i nterleukin-6, NO: nitric oxide; SOD: superoxide dismutase; GPx: glutathione peroxidase; VCAM-1: vascular cell adhesion molecule-1; HUVECs: human umbilical vein endothelial cells; TC: total cholesterol; TG: triglycerides; VLDL-C: very low density lipoprotein cholesterol; HDL-C: high density lipoprotein cholesterol; LDL: low-density lipoprotein; CAT: catalase; LPO: lipid peroxidation; GSH: glutathione; GST: glutathione-S-transferase; AGEs: advanced glycation end products; ECM: extracellular matrix; TGF-β: transforming growth factor- β; MMPs: matrix metalloproteinases; cTnT: cardiac troponin T; LDH: lactate dehydrogenase; ROS: reactive oxygen species; LAP: leucine aminopeptidase; y-GTP ;y-glutamyl transpeptidase; Bcl-2: B-cell lymphoma 2; IL-21: interleukin-21; IL-23: interleukin-23; UDP-GT: UDP glucuronosyltransferase; NQO1: NAD(P)H quinone dehydrogenase-1; MPO: myeloperoxidase; ALT: alanine aminotransferase; AST: aspartate aminotransferase; ALP: alkaline phosphatase; CCl4: carbon tetrachloride, HA; hyaluronic acid; MDA: malondialdehyde; γ -GT: γ -glutamyl transferase; ALP: alkaline phosphatase; ALA-D: aminolevulinic acid dehydratase; PDX-1: pancreas/duodenum homeobox 1; PPAR-γ: peroxisome proliferator-activated receptor γ; TBARS: 2-thiobarbituric acid reactive substances; MFB: medial forebrain bundle; H2O2: hydrogen peroxide; MWM: Morris water maze; EPM: elevated plus maze; MPT: membrane permeability transition; LPS: lipopolysaccharide; BUN: blood urea nitrogen; AChE: acetyl cholinesterase; MAO-A: Monoamine oxidase-A; G6PD: glucose-6-phosphate dehydrogenase; MAPKs: mitogen-activated protein kinases; AEGs: Advanced glycation end products; GLUT1: glucose transporter protein 1, GLUT4: glucose transporter protein 4; PI3K: phosphatidylinositol 3-kinase; p-Akt: phosphorylated protein kinase B; TAG: triacylglycerol; GSSG: glutathione disulfide;, AUCglucose: area under the curve for glucose; HOMA-IR: homeostasis model assessment insulin resistance; IRS-1: insulin receptor substrate-1; IR: insulin receptor; GLUT-2: glucose transporter protein 2;; F-1,6-BP: fructose-1,6-bisphosphatase; PFK: phosphofructokinase; a-CN: a-casein; STZ: streptozotocin; β-LG: β-lactoglobulin; Aβ: amyloid β Protein; (Ca2+): cytosolic Ca2+ concentration; MTT: 3-(4,5-dimethylthiazole-2-yl)-2,5-diphenyl-tetrazolium bromide; H2DCF-DA: Hoechst 33342 dye, fluo-4 AM & 2_,7_-dichlorodihydrofluorescin diacetate; n-NOS: neuronal nitric oxide synthase; SNCV: sciatic nerve conduction velocity; LTP: long-term potentiation; PS Amp: population spikes amplitude; AUC: area under curve; CA-1: region I of hippocampus proper; AD: Alzheimer disease; PC12: pheochromocytoma cells; Bcl-2: B-cell lymphoma 2; JNK: c-Jun N-terminal protein kinase; , ICR: institute of cancer research; peroxidase, MDA: malondialdehyde, k-CN: kappa-casein, DCF: dichlorofluorescin, ; PGE2: prostaglandin E2; e-NOS: endothelial nitric oxide synthase; HSCs: hepatic stellate cells; UVB: ultraviolet B; TAC: total antioxidant capacity; L-NAME: NG-nitro-L-argininemethyl ester; SBP: systolic blood pressure; LV: left ventricle; HDAC: histone deacetylase; VEGF: vascular endothelial growth factor.