Symbol	Definition	Value	Symbol	Definition	Value
τmPC	PC membrane time constant	50 ms	NPC	Number of PCs in recurrent network	3000 (Figures 2–3) 12000 (Figure 4)
EPCleak	PC leak potential	−68 mV	NINH	Number of INHs in recurrent network	300 (Figures 2–3) 1200 (Figure 4)
vPCth	PC spike threshold	−36 mV	wPC,G	Gating input weight without LTP-IE	0.8216 (Figures 2–3) 0.5 (Figure 4)
τPCr	PC refractory period	8 ms	wPC,PC	Excitatory recurrent weight scale factor	2.6
τmINH	INH membrane time constant	5 ms	wminPC,PC	Min nonzero recurrent weight	.1
EPCleak	INH leak potential	−60 mV	wPC,INH	INH→PC connection weight	0.02 (Figures 2–3) 0.002 (Figure 4)
vPCth	INH spike threshold	−50 mV	wINH,PC	PC→INH connection weight	.03
τPCr	INH refractory period	2 ms	pPC,INH	INH→PC connection probability	0.5
EE	Excitatory synaptic reversal potential	0 mV	pINH,PC	PC→INH connection probability	0.5
τE	Excitatory synaptic time constant	2 ms	rG	Gating input firing rate	125 Hz (Figures 2–3) 200 Hz (Figure 4)
EI	Inhibitory synaptic reversal potential	−80 mV	λPC,PC	Excitatory recurrent connectivity length scale	.053 m
τI	Inhibitory synaptic time constant	2 ms	σmax	Max LTP-IE value	2 (Figures 2–3) 1.84 (Figure 4)
rmax	Max position-driven firing rate	20 Hz	rσ	Threshold firing rate for LTP-IE	10 Hz
λPL	Position-tuning length constant	0.15 m	βσ	Scale factor for LTP-IE onset	1
			ΔT	Simulation time step	0.5 ms