Example rFN neuron with body-centered tuning. A–C, Contour plots illustrating the cell's 3D spatial tuning for translation with the head upright (A) as well as after vertical plane head reorientation 30° toward right-ear-down (B) and after horizontal plane reorientation 45° to the left (C). Star: PD; pink circle: predicted PD for head-centered tuning; blue square: predicted PD for body-centered tuning. D, STC model fit to gains and phases across elevations for an azimuth angle of 45° (i.e., along dashed white line in B) with the head upright (blue lines/symbols) and after reorientation toward ear-down (red lines/symbols). E, STC model fit to gains and phases across azimuth angles for 0° elevation (i.e., along dashed white line in C) with the head upright (blue lines/symbols) and after reorientation to the left (green lines/symbols). Dashed black curves in D and E show the predictions for head-centered tuning.