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. 2019 Jun 20;10:773. doi: 10.3389/fpls.2019.00773

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Copyright © 2019 Hesse, Zelkowski, Mikhailova, Keijzer, Houben and Schubert.

This is an open-access article distributed under the terms of the Creative Commons Attribution License (CC BY). The use, distribution or reproduction in other forums is permitted, provided the original author(s) and the copyright owner(s) are credited and that the original publication in this journal is cited, in accordance with accepted academic practice. No use, distribution or reproduction is permitted which does not comply with these terms.

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SEM imaging reveals the ultrastructure of rye bivalents during prophase I. The images of the right column show the regions of interest enlarged. (A) Top view of aligned homologous chromosomes inside a meiocyte at zygotene. Chromatin clusters (chromomeres) are clearly visible at the chromosome surface (arrow). (B) Cross section of a bivalent inside a meiocyte during zygotene-pachytene. The bivalent in the green rectangle is composed of two paired homologs (blue arrows) both containing two chromatids. The red arrow indicates the SC.