Asymmetric units of GII.2 T = 1, GII.2 T = 3, and GII.4 T = 4 particles. The icosahedral symmetry axes positions are indicated by a pentagon (fivefold), triangle (threefold), and oval (twofold). (A) Shell domain of a single subunit in each ASU of GII.2 T = 1 particles. (B) Shell domains of three subunits in quasi-equivalent positions, A, B, and C, that constitute each ASU of GII.2 T = 3 particles. (C) GII.4 shell domains in the T = 4 particle ASU with four subunits, A, B, C, and D. The C–D dimer takes up a position similar to the C–C dimer in T = 3 particles.