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| Description of variable | Symbol | Dimension | Constraints on value |
|---|---|---|---|
| Vector of mutation sign indicators | μ | 1‥nm | Integer, 0‥1 |
| Matrix of mutation effects | E | [0‥1] × [1‥nm] | Positive real number |
| Vector of frequencies of negative and positive-effect mutations | q | 0‥1 | 0 < q0 < 1 |
| Vector of shape parameters | α | 0‥1 | Positive real number |
| Vector of scale parameters | β | 0‥1 | Positive real number |
