1	Black spades on the hind legs	2
–	Spades of light coloration	3
2	Large body (6–9 cm) without orange dots, spades entirely black; Spain, Portugal and southern France	P. cultripes
–	Small body (<6 cm) with orange dots, spades bordered with black; Morocco	P. varaldii
3	Domed skull, developed webbing, and small body (<6 cm)	4
–	Flat skull, partial webbing, and large body (6–8 cm)	5
4	Dorsum stripes rare; Central and northwestern Europe, west of a Crimea–Moscow imaginary line	P. fuscus
–	Three dorsum stripes often present; Eastern Europe and Central Asia, east of a Crimea–Moscow imaginary line	P. vespertinus
5	Levantine region (Israel, Lebanon, and Syria)	P. syriacus syriacus
–	Caucasus and Caspian Sea shores, Anatolia, and European Turkey	P. syriacus boettgeri
–	Balkan Peninsula, except Peloponnese	P. balcanicus balcanicus
–	Peloponnese	P. balcanicus chloeae