Potential effects of sensory conflict (scene tilt) applied during the movement of the head. In the memorization phase (left), subjects evaluated the orientation of the target (green beams). If a 24/15 gain between the actual and virtual head roll motion was applied when the subject tilted the head by 15° to the left (right), the gravitational and the visual vertical no longer matched and the visual scene was tilted 9° (24/15 × 15° − 15° = 9°) with respect to gravity in the opposite direction of the head tilt. If subjects used the memorized target angle with respect to the visual background, their responses would be deviated 9° in reproducing the earth-fixed target orientation. Conversely, if the target orientation was memorized and reproduced with respect to the body, gravity, or the physical world, no systematic deviation would occur.