Dynamics of collective cell migration in heterogeneous tissue. (Top) The time course for cell positions (front (red) , center (black) , and back (blue) for ) are shown for different trailing cell properties, relative to the lead cell, with (A) low β, (B) high β, and (C) low δ trailing cells. See the text for details. (Bottom) The spatial pattern of junctional force averages as a function of the cell-cell junction is shown. Parameters are lead cell: δ = 0.9, β = 0.16. Trailing cells have a scaling factor of 0.2: (A) δ = 0.9, β = 0.1082, (B) δ = 0.9, β = 0.2454, and (C) δ = 0.1885, β = 0.16. To see this figure in color, go online.