Table 1.
Gesture type (gestural action) | Definition | N | Chimpanzee | ||
---|---|---|---|---|---|
I | J | All | |||
Arm benda | One or both arms held horizontally away from the signaller and then rapidly retracted towards the signaller | 10 | − | − | − |
Arm raise | Raise arm and/or hand vertically in the air | 56 | + | + | + |
Arm raise w. object | As ‘Arm raise’, while holding object | 10 | − | + | + |
Arm shake | Small repeated back and forth motion of the arm, and/or hand | 5 | + | + | + |
Arm wave | Large repeated back and forth motion of the arm raised above the shoulder | 7 | − | + | + |
Arm wave w. object | As ‘Arm wave’, while holding object | 5 | − | + | + |
Bite | Recipient’s body is held between the teeth or lips of the signaller | 2 | + | + | + |
Clap | Both palms moved towards each other and are brought together with audible contact | 6 | 1 | − | 1 |
Embrace | Signaller wraps one or both arm(s) around the recipient and maintains physical contact | 3 | + | + | + |
Fling | Rapid movement of the arm or hand in the direction of the recipient | 1 | 1 | + | + |
Grab | Hand is firmly closed over part of the recipient’s body | 16 | + | + | + |
Grab hold | As ‘Grab’ but closed hand contact is maintained for at least 2 s | 11 | + | + | + |
Grab pull | As ‘Grab’ but closed hand contact is maintained and a force exerted to move the recipient from their current position | 8 | + | + | + |
Grab pull 2hands | As ‘Grab pull’ but with both hands | 1 | + | + | + |
Hand on | Palm of the hand or knuckles are placed on the recipient, contact lasts for more than 2 s | 1 | + | + | + |
Hand wavea | Repeated back and forth motion of the hand from the wrist, typically while held above the shoulder | 21 | − | − | − |
Head shake | Repeated back and forth movement of the head (side to side or vertical) | 7 | − | + | + |
Hit object/ground | Movement of the arm from the shoulder with hard short contact of the open palm or closed fist to an object or the ground | 10 | + | + | + |
Hit object/ground 2hands | As ‘Hit object/ground’ but with both hands | 2 | + | + | + |
Hit object w. object | As ‘Hit object/ground’ but the hand holds an object which is brought into contact with another object or the ground | 5 | − | + | + |
Hit other | As ‘Hit object/ground’ but the hand is brought into contact with the recipient’s body | 17 | + | + | + |
Hit other 2hands | As ‘Hit other’ but with both hands | 3 | + | + | + |
Hit selfb | As ‘Hit other’ but the hand is brought into audible contact with the signaller’s body | 10 | − | 1 | 1 |
Hit with object | As ‘Hit other’ but the hand holds an object which is brought into contact with the recipient’s body | 3 | + | + | + |
Hitting object/ground | As ‘Hit object/ground’ but there is regular rhythmic repetition of the action | 8 | 1 | 1 | + |
Hitting other | As ‘Hit other’ but there is regular rhythmic repetition of the action | 5 | + | + | + |
Hitting with object | As ‘Hit with object’ but there is regular rhythmic repetition of the action | 1 | − | − | + |
Jump | While bipedal both feet leave the ground simultaneously, accompanied by horizontal displacement through the air | 2 | + | + | + |
Locomote | An exaggerated stiff walking or running movement, typically with audible contact of the feet | 2 | + | + | + |
Look | Signaller holds eye-contact position with the recipient for at least 2 s | 3 | 1 | 1 | + |
Object in mouth | Signaller approaches recipient while carrying an object in the mouth | 2 | + | + | + |
Object on headb | Signaller faces or approaches recipient while balancing an object on the head | 4 | − | 1 | 1 |
Object move | Object is displaced in one direction, contact is maintained throughout movement | 2 | + | + | + |
Object shake | Repeated back and forth movement of an object | 10 | + | + | + |
Poke | Firm, brief push of one or more fingers into the recipient’s body | 1 | 1 | + | + |
Push | Palm in contact with recipient’s body and force is exerted in attempt to displace recipient | 7 | + | + | + |
Reach palm | Arm extended to the recipient with the palm held vertically or upwards and the fingers in an open position | 107 | + | + | + |
Reach directed | As ‘Reach palm’ but arm is extended towards a third party or object, while audience checking, response waiting, and/or other signals are directed to recipient | 164 | − | + | + |
Rocking | Back and forth movement of the torso from the waist, typically while seated | 6 | 1 | + | + |
Rubb | Back and forth movement of the palms on the signaller’s body | 32 | − | − | − |
Stomp | Sole of the foot is lifted vertically and brought into contact with the surface being stood upon | 11 | + | + | + |
Stomping | As ‘Stomp’ but there is regular rhythmic repetition of the action | 1 | + | + | + |
Swing | Large back and forth movement of arm(s) or leg(s) from shoulder or hip | 3 | + | + | + |
Swing w. object | As ‘Swing’ but hand or foot holds an object | 7 | − | + | + |
Tap object | Movement of the arm from the wrist or elbow, with firm short contact of one or more fingers to the object | 2 | − | − | + |
Tap other | As ‘Tap object’ but the fingers are brought into contact with the recipient’s body | 3 | + | + | + |
Tapping object | As ‘Tap object’ but there is regular rhythmic repetition of the action | 6 | − | 1 | + |
Tapping other | As ‘Tap other’ but there is regular rhythmic repetition of the action | 6 | − | + | + |
Tapping selfb | As ‘Tapping other’ but the fingers are brought into contact with the signaller’s body | 3 | − | − | − |
Throw object | Object is moved and released so that there is displacement through the air after moment of release | 36 | + | + | + |
Thrust | Hips are brought into repeated contact with the recipient’s body | 1 | − | − | + |
Touch | Light contact of the hand and/or fingers on the body of the recipient, contact under 2 s | 25 | + | + | + |
Within the chimpanzee repertoire we distinguish those recorded to date as employed by infant (I) or juvenile (J) chimpanzees, from those recorded in the species repertoire (All); + = present, − = not observed; 1observed but insufficient cases in wild chimpanzees for inclusion in current repertoire. Gesture descriptions follow: Hobaiter and Byrne 2011, updated in Hobaiter and Byrne 2017. N = number of gesture tokens. Given a small data set, all potential gestures are described here, however we would typically require n = 2 cases of intentional use and use by at least two individuals for inclusion in a great ape repertoire (Hobaiter and Byrne 2011a, b)
aThe gestures have not been described to date in the repertoire of other apes
bThe gesture is found in the gorilla, but not in the chimpanzee repertoire; all other gestures are present in the chimpanzee repertoire