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Least square means (± SEM) of heart rate variation measured at rest, during loading, during transport, during unloading and on the return to the home pen. Least square means without a common superscript differ (Tukey’s test; P < 0.001). GH1: One lap of the handling course with gentle handling; RH1: One lap of the handling course with rough handling; GH3: Three laps of the handling course with gentle handling; RH3: Three laps of the handling course with rough handling.
