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. 2019 Aug 7;9:11440. doi: 10.1038/s41598-019-47725-0

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© The Author(s) 2019

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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The cartoon representation of membrane-embedded (a) wild-type lysenin in a pure POPC lipid bilayer used in MD simulations. E84A and E85A residues are shown in the van der Waals surface representation (colored red). (b) E84K-E85K lysenin mutant in a mixed lipid bilayer consisting of POPC (orange), POPI (blue), POPE (magenta) CHOL (yellow), and SM (green). E84K and E85K residues are shown in the van der Waals surface representation (colored blue).