Table 5.
Descriptive statistics for morphometric variables of Pristimantis, subgenus Huicundomantis. Mean ± SD followed by the range of the measurements are given in each cell. ‘n’ is for the number of samples. Numbers in brackets after the species name refer to the bibliographic source of the data: (1) Lynch 1979, (2) Bustamante and Mendelson 2008, (3) Yánez-Muñoz et al. 2016, (4) Duellman and Pramuk 1999, (5) Lynch and Duellman 1995, (6) Brito et al. 2016; otherwise the source is this study. Abbreviations are: SVL = snout-vent length; HL = head length; HW = head width; TD = tympanum diameter; ED = eye diameter. All measurements are in mm.
| Females | |||||
|---|---|---|---|---|---|
| Clade | SVL | HL | HW | TD | ED |
| P. atillo sp. nov. n = 5 | 31.3 ± 2.3; 29.4–35.3 | 11.0 ± 0.5; 10.3–11.7 | 11.5 ± 0.7; 10.9–12.7 | 1.7 ± 0.2; 1.5–1.8 | 3.2 ± 0.1; 3.1–3.3 |
| P. atratus (1) n = 10 | 27.4 ± 2.1; 24.9–29.2 | – | – | – | – |
| P. balionotus (1) n = 7 | 28.1 ± 0.6; 27.1–29.1 | – | – | – | – |
| P. cryptomelas (1) n = 1 | 38.6 | – | – | – | – |
| P. gagliardoi (2) n = 5 | 30.6 ± 3.1; 26.8–33.6 | 10.7 ± 0.6; 9.8–11.4 | 12.2 ± 1.0; 10.7–13.3 | 1.3 ± 0.2; 1.0–1.5 | 3.8 ± 0.3; 3.5–4.1 |
| P. gloria sp. nov. n = 15 | 30.1 ± 3.0; 26.7–35.8 | 11.0 ± 0.7; 10.1–12.4 | 11.3 ± 1.1; 9.8–13.5 | 1.6 ± 0.1; 1.4–1.8 | 3.2 ± 0.2; 2.8–3.6 |
| P. hampatusami (3) n = 11 | 30.6 ± 2.1; 25.9–34.1 | – | – | – | – |
| P. jimenezi sp. nov. n = 9 | 35.0 ± 2.1; 31.1–37.4 | 12.0 ± 0.5; 11.0–12.6 | 13.1 ± 0.6; 13.0–13.9 | 1.9 ± 0.1; 1.7–2.1 | 3.7 ± 0.2; 3.5–4.0 |
| P. lutzae sp. nov. n = 15 | 31.4 ± 1.3; 29.7–33.9 | 11.3 ± 0.4; 10.7–12.1 | 12.2 ± 0.5; 11.4–12.9 | 1.7 ± 0.1; 1.5–1.8 | 3.4 ± 0.2; 3.1–3.8 |
| P. multicolor sp. nov. n = 10 | 35.3 ± 3.5; 29.4–40.5 | 13.4 ± 1.1; 11.6–15.1 | 14.4 ± 1.3; 11.9–16.1 | 2.2 ± 0.2; 1.8–2.5 | 4.0 ± 0.3; 3.6–4.4 |
| P. muscosus (4) n = 4 | 37.8; 29.6–46.1 | – | – | – | – |
| P. nangaritza sp. nov. n = 4 | 29.1 ± 2.7; 25.9–32.4 | 11.5 ± 0.8; 10.6–12.2 | 11.2 ± 1.0; 10.1–12.1 | 1.5 ± 0.1; 1.4–1.7 | 3.7 ± 0.3; 3.4–4.0 |
| P. percultus (1) n = 1 | 38.2 | – | – | – | – |
| P. phillipi (5) n = 6 | 31.2 ± 1.1; 26.5–33.7 | – | – | – | – |
| P. phoxocephalusn = 5 | 36.9 ± 2.2; 34.2–39.9 | 12.9 ± 0.9; 11.5–13.8 | 13.1 ± 1.2; 11.3–14.2 | 1.9 ± 0.1; 1.8–2.1 | 3.8 ± 0.3; 3.5–4.1 |
| Pristimantis CCS1 n = 1 | 37.0 | 12.5 | 13.8 | 2.0 | 3.5 |
| Pristimantis CCS2 n = 1 | 38.3 | 13 | 14.4 | 1.7 | 3.5 |
| Pristimantis UCS1 n = 1 | 31.2 | 12.4 | 12.4 | 1.7 | 3.8 |
| P. spinosus (1) n = 29 | 31.8 ± 1.62; 28.3–34.5 | – | – | – | – |
| P. tinguichaca (6) n = 9 | 29.7 ± 1.5; 28.1–31.7 | 10.6 ± 0.4; 10.1–11.0 | 11.0 ± 0.5; 10.1–11.5 | 1.4 ± 0.1; 1.3–1.7 | 3.3 ± 0.3; 2.8–3.7 |
| P. torresi sp. nov. n = 5 | 34.7 ± 3.7; 30.1–39.5 | 12.2 ± 1.3 10.4–13.8 | 13.1 ± 1.2 11.5–14.6 | 1.9 ± 0.2 1.8–2.2 | 3.6 ± 0.3 3.3–3.9 |
| P. totoroi sp. nov. n = 7 | 33.0 ± 0.9; 31.9–34.3 | 12.1 ± 0.4; 11.5–12.6 | 12.3 ± 0.3; 11.8–12.6 | 1.6 ± 0.1; 1.6–1.8 | 3.3 ± 0.2; 3.1–3.6 |
| P. verrucolatus sp. nov. n = 2 | 43.6 ± 4.5; 40.4–46.8 | 15.0 ± 1.1; 14.2–15.8 | 16.6 ± 1.7; 15.3–17.8 | 2.2 ± 0.1; 2.1–2.3 | 4.2 ± 0.0; 4.2–4.3 |
| P. versicolorn = 4 | 27.8 ± 3.5; 24.9–32.4 | 10.7 ± 1.6; 9.0–12.4 | 10.5 ± 1.0; 9.5–11.4 | 1.7 ± 0.3; 1.4–2.1 | 3.2 ± 0.4; 2.9–3.7 |
| Males | |||||
| P. atillo sp. nov. n = 29 | 24.7 ± 2.5; 17.7–28.1 | 8.7 ± 0.7; 6.4–9.7 | 8.8 ± 0.9; 5.9–10.0 | 1.2 ± 0.1; 0.8–1.4 | 2.7 ± 0.3; 2.0–3.0 |
| P. atratus (1) n = 19 | 21.7 ± 3.71; 17.4–24.0 | – | – | – | – |
| P. balionotus (1) n = 2 | 20.0 ± 0.2; 21.8–22.2 | – | – | – | – |
| P. chomskyi sp. nov. n = 3 | 28.0 ± 4.2; 24.0–32.4 | 9.7 ± 1.2; 8.8–11.1 | 10.6 ± 1.6; 9.4–12.4 | 1.3 ± 0.2; 1.1–1.5 | 3.3 ± 0.4; 2.9–3.6 |
| P. cryptomelas (1) n = 4 | 29.2; 28.2–30.3 | – | – | – | – |
| P. gagliardoi (2) n = 5 | 22.2 ± 2.0; 19.1–24.3 | 7.9 ± 0.9; 6.8–9.1 | 9.0 ± 0.9; 7.7–10.2 | 0.8 ± 0.0; 0.7–0.8 | 2.8 ± 0.2; 2.4–3.0 |
| P. gloria sp. nov. n = 24 | 21.7 ± 2.3; 16.8–24.7 | 8.6 ± 0.7; 6.7–9.4 | 8.4 ± 0.8; 6.5–9.5 | 1.2 ± 0.1; 0.9–1.4 | 2.6 ± 0.2; 2.1–2.9 |
| P. hampatusami (3) n = 28 | 21.0 ± 1.8; 17.0–24.9 | – | – | – | – |
| P. jimenezi sp. nov. n = 12 | 25.5 ± 1.6; 21.8–27.1 | 8.9 ± 0.6; 7.8–9.6 | 9.1 ± 0.6; 7.8–9.7 | 1.4 ± 0.1; 1.3–1.5 | 3.06 ± 0.24; 2.56–3.37 |
| P. lutzae sp. nov. n = 14 | 24.6 ± 1.7; 21.4–27.0 | 8.7 ± 0.5; 7.9–9.4 | 9.3 ± 0.7; 8.2–10.4 | 1.3 ± 0.1; 1.1–1.4 | 2.77 ± 0.19 2.53–3.26 |
| P. multicolor sp. nov. n = 12 | 26.2 ± 3.5; 19.7–29.7 | 9.6 ± 1.0; 8.0–11.0 | 10.1 ± 1.2; 8.0–11.5 | 1.5 ± 1.2; 1.1–1.8 | 3.18 ± 0.49; 2.45–3.82 |
| P. nangaritza sp. nov. n = 13 | 18.8 ± 1.1; 17.4–20.8 | 7.6 ± 0.6; 7.1–8.8 | 7.2 ± 0.5; 6.6–8.3 | 1.0 ± 0.1; 0.8–1.2 | 2.69 ± 0.17 2.49–3.08 |
| P. percultus (1) n = 1 | 29.8 | – | – | – | – |
| P. phillipi (5) n = 10 | 23.0 ± 0.4; 21.1–25.1 | – | – | – | – |
| P. phoxocephalusn = 3 | 24.7 ± 3.2; 20.8–27.9 | 9.0 ± 1.36; 7.4–10.6 | 8.7 ± 1.5; 7.0–10.4 | 1.2 ± 0.2; 1.0–1.3 | 2.66 ± 0.25 2.36–2.90 |
| Pristimantis UCS2 n = 2 | 20.5 ± 1.7; 19.3–21.7 | 7.6 ± 0.7; 7.1–8.1 | 7.3 ± 0.4; 7.0–7.6 | 1.0 ± 0.1; 0.9–1.0 | 2.43 ± 0.23 2.29–2.62 |
| P. spinosus (1) n = 34 | 20.1 ± 1.8; 16.1–25.0 | – | – | – | – |
| P. teslai sp. nov. n = 4 | 25.2 ± 1.8; 23.4–27.3 | 8.9 ± 0.4; 8.4–9.2 | 9.0 ± 0.6; 8.2–9.5 | 1.3 ± 0.1; 1.2–1.5 | 2.89 ± 0.14 2.76–3.07 |
| P. tinguichaca (6) n = 11 | 23.4 ± 1.1; 21.2–24.7 | 8.7 ± 0.3; 8.2–9.2 | 8.6 ± 0.5; 7.9–9.4 | 1.3 ± 0.2; 1.1–1.6 | 2.8 ± 0.4 2.3–3.3 |
| P. torresi sp. nov. n = 18 | 25.9 ± 2.1; 23.3–30.0 | 9.2 ± 0.6; 8.4–10.5 | 9.3 ± 0.7; 8.4–10.5 | 1.3 ± 0.1; 1.2–1.6 | 2.98 ± 0.23 2.64–3.47 |
| P. totoroi sp. nov. n = 21 | 26.8 ± 2.0; 23.2–29.4 | 9.6 ± 0.6; 8.5–10.5 | 9.5 ± 0.6; 8.4–10.3 | 1.3 ± 0.1; 1.1–1.4 | 2.79 ± 0.25 2.27–3.25 |
| P. verrucolatus sp. nov. n = 15 | 29.4 ± 2.7; 25.1–34.5 | 9.7 ± 0.7; 8.5–10.7 | 10.5 ± 1.0; 8.5–12.1 | 1.5 ± 0.1; 1.3–1.8 | 3.21 ± 0.24 2.79–3.64 |
| P. versicolorn = 12 | 20.5 ± 1.6; 18.1–23.3 | 8.2 ± 0.5; 7.3–8.9 | 7.6 ± 0.6; 6.7–8.6 | 1.3 ± 0.2; 1.0–1.6 | 2.84 ± 0.32 2.32–3.36 |