Table 1. Morphometric data of examined specimens of Hylobatrachus (all in mm).
For abbreviations of measurements, see Methods. Additional abbreviations: M, male; F, female, HT, holotype, ST, syntype. The column 16S indicates individuals for which a fragment of the 16S rRNA gene has been sequenced and included in the phylogenetic tree (Fig 1). Since the femoral glands in Hylobatrachus are usually poorly recognizable they were excluded from the measurements. The sex of the individuals was determined examination of femoral glands or by incision.
| Voucher number | Field number (tissue number) |
Sex | 16S | Location | SVL | HW | HL | TD | ED | END | NSD | NND | FORL | HAL | HIL | FOTL | FOL | TIBL |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| M. lugubris | ||||||||||||||||||
| MNHN 1994.1750 (ST) | — | F | Madagascar | 38.0 | 11.6 | 13.7 | 2.9 | 4.8 | 3.1 | 1.8 | 3.4 | 21.7 | 10.7 | 59.7 | 26.4 | 18.3 | 17.5 | |
| MNHN 1994.1751 (ST) | — | M | Madagascar | 31.5 | 10.3 | 11.9 | 3.6 | 4.3 | 3.0 | 1.3 | 3.3 | 19.0 | 9.4 | 49.4 | 22.5 | 15.6 | 15.1 | |
| MNHN 1994.1752 (ST) | — | M | Madagascar | 32.0 | 10.3 | 11.5 | 3.7 | 4.1 | 3.0 | 1.7 | 2.8 | 19.0 | 9.4 | 51.6 | 23.8 | 16.3 | 15.6 | |
| MNHN 4583 (ST) | — | M | Madagascar | 32.0 | 11.2 | 11.5 | 3.9 | 4.3 | 3.0 | 1.6 | 3.8 | 21.1 | 9.8 | 54.3 | 25.6 | 17.6 | 16.4 | |
| ZSM 166/2002 | MV 2001.1101 | F | + | Mantadia | 35.7 | 10.9 | 12.7 | 3.2 | 4.5 | 3.5 | 2.0 | 3.7 | 22.4 | 11.2 | 59.9 | 28.3 | 19.7 | 17.9 |
| ZSM 167/2002 | MV 2001.1102 | M | + | Mantadia | 33.6 | 11.3 | 13.0 | 4.1 | 4.5 | 3.3 | 1.8 | 3.1 | 21.4 | 10.6 | 54.8 | 26.4 | 18.1 | 16.8 |
| ZSM 750/2009 | ZCMV 7206 | M | + | Ambatoroma | 33.6 | 11.0 | 12.7 | 3.8 | 4.8 | 3.5 | 1.8 | 3.6 | 19.4 | 9.6 | 48.2 | 23.2 | 16.4 | 15.2 |
| ZSM 749/2009 | ZCMV 7205 | M | Ambatoroma | 31.3 | 10.3 | 11.8 | 3.7 | 4.6 | 2.9 | 1.9 | 3.6 | 19.3 | 9.5 | 50.4 | 23.1 | 16.1 | 15.2 | |
| ZSM 751/2009 | ZCMV 7236 | M | + | Ambatoroma | 31.9 | 11.2 | 12.0 | 3.4 | 4.8 | 2.8 | 2.4 | 3.8 | 19.5 | 9.1 | 50.0 | 21.5 | 13.8 | 15.6 |
| ZSM 747/2009 | ZCMV 11124 | F | + | Ambodirafia | 35.8 | 13.3 | 13.7 | 3.5 | 5.3 | 3.0 | 2.1 | 3.5 | 22.2 | 10.8 | 58.3 | 26.8 | 18.8 | 17.6 |
| ZSM 748/2009 | ZCMV 11129 | F | + | Ambodirafia | 36.8 | 12.0 | 13.0 | 3.0 | 5.4 | 3.4 | 1.7 | 3.9 | 22.6 | 10.8 | 56.1 | 25.7 | 18.1 | 17.1 |
| ZSM 65/2002 | MV 2001.1364 (2002-H39) |
Mantadia | 36.0 | 13.0 | 14.7 | 3.8 | 4.7 | 3.5 | 2.2 | 3.9 | NM | 10.7 | NM | NM | 18.3 | 18.9 | ||
| ZSM 299/2005 | FGZC 2668 | F | Vohidrazana | 36.2 | 12.7 | 14.2 | 3.6 | 5.4 | 3.8 | 2.2 | 3.7 | 24.1 | 11.2 | 56.3 | 28.5 | 19.3 | 17.8 | |
| ZSM 300/2005 | FGZC 2670 | F | Vohidrazana | 36.8 | 12.1 | 14.0 | 3.9 | 5.5 | 3.3 | 2.2 | 4.1 | 21.3 | 11.0 | 56.1 | 27.4 | 17.4 | 17.6 | |
| M. cowanii | ||||||||||||||||||
| ZSM 63/2002 | MV 2001.1353 (2002-H28) |
F | Mantadia | 43.5 | 14.1 | 15.6 | 2.9 | 5.3 | 3.9 | 2.7 | 4.1 | 28.5 | 12.9 | 67.1 | 28.6 | 17.8 | 22.3 | |
| ZSM 64/2002 | MV 2001.1365 (2002-H40) |
M | Mantadia | 32.9 | 11.2 | 12.8 | 3.7 | 4.1 | 3.1 | 1.9 | 2.7 | 21.0 | 8.8 | 54.3 | 24.1 | 12.7 | 16.6 | |
| ZSM 306/2005 | FGZC 2121 | F | + | Ambohitantely | 41.3 | 13.6 | 14.8 | 3.3 | 5.2 | 3.8 | 2.1 | 3.9 | 23.2 | 11.8 | 62.4 | 28.8 | 19.5 | 20.2 |
| ZSM 301/2005 | FGZC 2673 | M | Vohidrazana | 35.7 | 12.2 | 13.5 | 3.3 | 4.9 | 3.6 | 2.3 | 4.3 | 23.1 | 9.8 | 60.8 | 27.7 | 18.1 | 19.5 | |
| ZSM 302/2005 | FGZC 2674 | M | Vohidrazana | 34.2 | 11.9 | 12.2 | 3.9 | 4.2 | 2.9 | 2.2 | 3.6 | 21.1 | 9.2 | 55.3 | 26.2 | 17.1 | 18.2 | |
| ZSM 171/2002 | MV 2001.1103 (2002-F29) |
M | + | Mantadia | 40.4 | 13.7 | 14.3 | 3.2 | 5.1 | 3.5 | 2.5 | 4.0 | 25.3 | 12.5 | 68.4 | 31.5 | 21.6 | 20.7 |
| ZSM 297/2005 | FGZC 2155 | F | Ambohitantely | 38.7 | 12.8 | 13.1 | 2.9 | 4.2 | 3.2 | 2.1 | 3.4 | 22.9 | 11.8 | 62.5 | 29.1 | 20.2 | 19.7 | |
| ZSM 298/2005 | FGZC 2158 | F | Ambohitantely | 39.9 | 13.1 | 13.5 | 2.8 | 4.3 | 3.3 | 2.6 | 4.0 | 22.1 | 11.1 | 64.9 | 29.5 | 20.7 | 19.7 | |
| ZSM 296/2005 | FGZC 2154 | M | Ambohitantely | 31.9 | 11.1 | 11.5 | 2.9 | 4.3 | 3.2 | 2.0 | 3.8 | 19.8 | 9.3 | 54.0 | 23.8 | 16.4 | 16.1 | |
| M. petakorona sp. nov. | ||||||||||||||||||
| ZSM 305/2005 | FGZC 2767 | F? | + | Marojejy | 29.0 | 10.1 | 10.8 | 2.6 | 3.5 | 2.8 | 1.5 | 3.1 | 19.3 | 8.9 | 53.9 | 25.3 | 14.9 | 17.1 |
| ZSM 501/2016 | ZCMV 15100 | M | Marojejy | 31.3 | 10.0 | 11.6 | 3.7 | 4.9 | 3.0 | 1.8 | 3.1 | 19.6 | 8.8 | 52.8 | 24.2 | 14.7 | 16.1 | |
| ZSM 502/2016 | ZCMV 15104 | F | + | Marojejy | 34.0 | 10.1 | 11.7 | 2.9 | 4.9 | 2.8 | 1.7 | 3.1 | 20.4 | 8.2 | 58.2 | 26.0 | 17.8 | 16.9 |
| ZSM 503/2016 | ZCMV 15106 | M | + | Marojejy | 28.9 | 9.2 | 10.6 | 3.4 | 4.4 | 2.7 | 1.5 | 2.5 | 18.3 | 8.4 | 46.9 | 22.5 | 14.4 | 15.3 |
| ZSM 504/2016 (HT) | ZCMV 15110 | F | + | Marojejy | 34.0 | 11.4 | 12.0 | 2.8 | 6.3 | 2.6 | 2.1 | 3.3 | 20.7 | 9.7 | 57.7 | 26.2 | 16.5 | 18.3 |
| ZSM 505/2016 | ZCMV 15111 | F | + | Marojejy | 27.0 | 9.2 | 10.1 | 2.0 | 4.8 | 1.9 | 1.5 | 2.2 | 17.7 | 8.6 | 50.4 | 23.0 | 13.9 | 16.1 |
| M. atsimo sp. nov. | ||||||||||||||||||
| ZSM 149/2004 | FGZC 277 | F | + | Manantantely | 34.8 | 11.7 | 12.3 | 2.7 | 4.6 | 3.8 | 1.8 | 3.1 | 22.0 | 10.7 | 56.7 | 26.0 | 17.5 | 17.5 |
| ZSM 150/2004 | FGZC 281 | F | Manantantely | 33.7 | 11.9 | 12.5 | 2.4 | 4.6 | 3.8 | 2.1 | 3.5 | 20.1 | 10.0 | 50.6 | 24.0 | 16.9 | 15.9 | |
| ZSM 72/2004 | FGZC 122 | F | + | Andohahela | 34.6 | 11.9 | 12.3 | 2.6 | 4.3 | 3.7 | 1.6 | 2.9 | 21.0 | 10.1 | 56.2 | 25.0 | 16.9 | 16.9 |
| ZSM 69/2004 (HT) | FGZC 116 | F | + | Andohahela | 34.5 | 11.8 | 12.8 | 2.2 | 5.4 | 3.5 | 2.0 | 3.7 | 20.7 | 10.3 | 53.3 | 24.2 | 16.7 | 16.1 |
| ZSM 174/2002 | MV 2001.1483 (2002-B20 = B21) |
F | + | Pic St. Louis | 33.8 | 11.9 | 12.2 | 2.7 | 4.8 | 3.4 | 1.6 | 3.1 | 19.7 | 9.6 | 53.6 | 23.4 | 15.7 | 16.1 |
| ZSM 253/2002 | 2002-A95/A99/B7 | F | + | Nahampoana | 31.2 | 10.5 | 11.4 | 2.2 | 4.3 | 3.4 | 1.6 | 3.0 | 18.6 | 8.9 | 52.2 | 23.3 | 15.5 | 16.6 |
| ZSM 172/2002 | MV 2001.1476 (2002-A95/A99/B7) |
F | + | Nahampoana | 34.1 | 11.7 | 12.6 | 2.2 | 4.9 | 3.2 | 1.7 | 2.8 | 21.0 | 9.6 | 56.1 | 24.7 | 16.6 | 16.6 |
| ZSM 367/2016 | ZCMV14843 | M | + | Anosy Massif | 25.2 | 9.9 | 11.4 | 3.0 | 5.2 | 3.0 | 1.8 | 3.0 | 17.4 | 8.7 | 46.3 | 21.2 | 11.0 | 15.0 |
| ZSM 368/2016 | ZCMV14844 | M | + | Anosy Massif | 28.0 | 10.2 | 12.0 | 3.3 | 5.0 | 3.0 | 2.0 | 2.8 | 17.0 | 8.3 | 47.7 | 21.8 | 14.3 | 14.9 |
| M. sp. Ca48 | ||||||||||||||||||
| ZSM 493/2006 | ZCMV 2821 | M | Ranomafana | 31.5 | 10.4 | 12.5 | 3.6 | 4.5 | 2.9 | 1.9 | 2.5 | 19.0 | 9.0 | 49.0 | 22.9 | 14.7 | 16.5 | |
| ZSM 494/2006 | ZCMV 2832 | F | Ranomafana | 37.8 | 13.1 | 14.8 | 2.7 | 5.6 | 3.2 | 2.5 | 2.5 | 24.9 | 10.7 | 62.7 | 28.5 | 18.9 | 18.6 | |
| ZSM 495/2006 | ZCMV 2856 | F | Ranomafana | 36.1 | 13.6 | 14.5 | 3.2 | 5.7 | 3.5 | 2.2 | 2.9 | 23.8 | 11.0 | 62.0 | 27.9 | 17.5 | 19.0 | |
| ZSM 496/2006 | ZCMV 2981 | M | Ranomafana | 26.9 | 10.2 | 11.8 | 3.1 | 4.5 | 2.1 | 1.5 | 3.3 | 19.4 | 8.9 | 52.2 | 23.1 | 13.6 | 15.7 | |
| ZSM 497/2006 | ZCMV 3046 | M | Ranomafana | 26.0 | 9.4 | 11.6 | 2.8 | 4.7 | 2.4 | 2.0 | 2.6 | 16.2 | 7.4 | 45.1 | 20.9 | 14.5 | 14.4 | |
| ZSM 498/2006 | ZCMV 3047 | F | Ranomafana | 38.1 | 15.2 | 14.8 | 2.9 | 4.7 | 3.6 | 2.2 | 2.9 | 22.4 | 11.0 | 58.2 | 27.2 | 18.4 | 17.8 | |
| ZSM 499/2006 | ZCMV 3118 | M | Ranomafana | 29.7 | 11.1 | 12.1 | 3.2 | 4.7 | 3.2 | 1.6 | 3.2 | 20.8 | 9.5 | 52.4 | 25.3 | 15.7 | 17.6 | |
| ZSM 717/2003 | FGMV 2002.0379 | ? | Ranomafana | 30.0 | 10.6 | 12.3 | 3.6 | 5.2 | 3.0 | 1.6 | 3.6 | 18.2 | 9.5 | 49.0 | 22.7 | 15.3 | 15.1 | |
| ZSM 718/2003 | FGMV 2002.0385 | F | Ranomafana | 36.5 | 12.2 | 12.7 | 4.1 | 5.0 | 2.9 | 2.3 | 4.6 | 21.0 | 10.3 | 58.7 | 27.2 | 15.1 | 17.7 | |
| ZSM 719/2003 | FGMV 2002.0386 | M | Ranomafana | 29.2 | 10.3 | 12.3 | 3.2 | 4.6 | 2.9 | 2.0 | 3.2 | 18.7 | 9.3 | 52.3 | 24.1 | 14.3 | 15.2 | |
| ZSM 720/2003 | FGMV 2002.0391 | F | + | Ambohitsara | 38.0 | 12.5 | 14.5 | 3.1 | 5.8 | 3.0 | 1.8 | 3.1 | 23.2 | 10.9 | 65.7 | 28.9 | 17.9 | 19.5 |
| ZSM 721/2003 | FGMV 2002.0392 | M | + | Ambohitsara | 28.6 | 10.9 | 12.7 | 3.5 | 5.0 | 2.9 | 2.0 | 3.2 | 19.4 | 8.8 | 50.6 | 23.1 | 15.0 | 16.1 |
| ZSM 722/2003 | FGMV 2002.0394 | M | Ranomafana | 30.1 | 10.9 | 12.0 | 2.4 | 4.8 | 3.2 | 2.1 | 4.5 | 20.6 | 9.4 | 57.2 | 26.1 | 16.7 | 17.7 | |
| ZSM 898/2006 | ZCMV 2813 | F | Ranomafana | 39.0 | 12.2 | 13.9 | 3.1 | 5.0 | 3.5 | 2.9 | 2.6 | 23.7 | 11.3 | 61.0 | 28.2 | 19.3 | 18.8 | |
| ZSM 2412/2007 | ZCMV 5933 | M? | Ambohitsara | 26.7 | 9.7 | 11.2 | 3.7 | 3.6 | 3.2 | 1.3 | 2.8 | 18.6 | 8.5 | 47.5 | 21.0 | 10.9 | 14.6 | |
| ZSM 730/2003 | FGMV 2002.0454 | M | Ranomafana | 26.8 | 10.3 | 11.8 | 2.8 | 5.0 | 2.9 | 1.8 | 3.2 | 19.3 | 7.9 | 48.7 | 23.2 | 13.2 | 19.5 | |
| ZSM 734/2003 | FGMV 2002.0460 | M | Ranomafana | 27.7 | 9.8 | 11.8 | 3.4 | 3.9 | 2.8 | 1.9 | 4.4 | 18.9 | 8.0 | 51.2 | 23.1 | 14.2 | 15.2 | |
| ZSM 646/2003 | FGMV 2002.181 | F | + | Ranomafana | 37.2 | 12.0 | 13.4 | 2.7 | 5.2 | 3.4 | 2.3 | 3.7 | 22.4 | 11.4 | 59.1 | 27.8 | 19.1 | 18.3 |
| ZSM 744/2001 | MV 2001.467 (2001-D21?) |
M | +? | Itremo | 26.8 | 8.9 | 9.5 | 2.2 | 3.9 | 2.1 | 1.6 | 2.4 | 16.6 | 8.1 | 44.9 | 20.8 | 13.5 | 13.7 |
| ZSM 745/2001 | MV 2001.468 (2001-D21?) |
M | +? | Itremo | 27.4 | 8.9 | 10.1 | 3.1 | 3.9 | 2.5 | 1.6 | 3.2 | 16.9 | 8.0 | 44.3 | 20.1 | 13.8 | 13.4 |