Table 2.

Measuring costs and benefits of key behaviours using movement

behavioural category	example issue	example metric	reference
optimal foraging theory	search behaviour	patch occupancy	[24]
trade-offs	dispersal decisions	net squared displacement	[25]
competition	costs to subordinates	rank-based space use	[26]
alternative strategies	male reproductive states	daily activity budget	[27]
group living	group size constraints	daily movement distance	[28]
parental investment	impact of maternal investment on survival	foraging trip length	[29]
behavioural syndromes	factors influencing different movement tactics	resource selection	[30]
plasticity	response to human activity	vagility	[8]
seasonal response	influence of longitudinal changes in resources	behavioural change point analysis	[31]