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Spatial randomness: mean fitness of mutants and wild-type cells at several consecutive time steps, starting from (a) m = 1 mutant, (b) m = 2 mutants, (c) m = 3 mutants and (d) m = 6 mutants. The total number of cells is N = 9. Simulations with random division rates are performed under the BD update rule on a circle, with σ = 0.9 and mutant and wild-type division rates anti-correlated; 10 000 realizations were used. (Online version in colour.)
