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. 2019 Sep 11;9:13176. doi: 10.1038/s41598-019-49059-3

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© The Author(s) 2019

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Phylogenetic trees based on the partial rrs gene sequences of Rickettsia (A), Anaplasma (B), Ehrlichia (C), and the TSA56 gene sequences of Orientia tsutsugamushi (D). All trees were mid-point rooted for clarity only. Bootstrap values (>70%) are shown for appropriate nodes. The scale bar represents number of nucleotide substitutions per site. The sequences from patient samples are marked in red, those from rodents and shrews are marked in blue, and sequences from ticks are marked in orange.