The coupling functions −Hinh(−φ), −Hexc(−φ), and −Hdesc(−φ), plotted as functions of φ, when only descending intersegmental connections are present.A, Schematic of a network containing descending-only intersegmental connections. B, The functions −Hexc(−φ) and −Hinh(−φ) generated numerically in a network containing a descending excitatory-only connection from cell 1A to 3B and inhibitory-only connection from cell 1B to 3A, respectively. The vertical dotted line marks the portion of the functions that sum to zero. C, The function −Hdesc(−φ) generated numerically in a network containing both excitatory and inhibitory connections as shown in A. When both descending connections are present, the stable phase lag occurs at ∼96° for the parameters of this representative simulation (arrow).