Table 1.

Model parameter description.

1-region parameter	Value	Description
Cl	135	Pyramidal to excitatory interneuron connection
C2	108	Excitatory interneuron to pyramidal connection
C3	33.75	Pyramidal to inhibitory interneuron connection
C4	33.75	Inhibitory interneuron to pyramidal connection
e	5	Maximum firing rate of a neuronal population
r	0.3	Steepness of sigmoid function
v0	6	Post-synaptic potential (PSP) for which 50% of e is achieved
A, B, kA * A	Estimated from data	Maximum PSP amplitude for pyramidal (A), inhibitory (B) and excitatory (kA * A) interneurons
a, b, ka * a	Estimated from data	Lumped representation of the sum of reciprocal of the membrane time constant and other spatially distributed delays in the dendritic network for pyramidal (a), inhibitory (b) and excitatory (ka * a) interneurons
2-region parameter
K1	Estimated from data	First to second region coupling strength
K2	Estimated from data	Feedback from 2nd region to the 1st region
Ad	Estimated from data	Inverse of delay time constant
P	Estimated from data	Constant added to external input p
External inputs
P	Gaussian (0,0.1)	Excitatory input from other cortical columns
I1	60 * unit impulse train	Electrical current input to pyramidal population
I2, I3	18 * unit impulse train	Electrical current inputs to interneurons