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Schematic of the assembly of prime-boost T4 nanoparticles. The DNA packaging machine is assembled by binding of gp17 motor at the portal of hoc−soc− phage T4 empty head (the cut-out of the head shows both the exterior and the interior) (A). Using the energy from ATP hydrolysis the motor packages DNA molecules into the head (B). Soc-fused antigens (C) and Hoc-tused targeting molecules (D) are then displayed on the heads.
