ndl1 is defective in reproductive organogenesis. (A–D) The ndl1 tassel phenotype is very variable, from mild (B) to severe (D). (E–H) SEMs of immature tassels in normal and mutant plants also show different expressivity of the ndl1 mutation. AMs, axillary meristems; IM, inflorescence meristem. (Scale bars, 500 μm.) (I–N) mRNA in situ hybridizations of immature tassels with ZYB15, ARF4, and BA1 antisense probes. (Scale bars, 250 μm.) (O and P) Confocal images of normal and ndl1 immature tassels expressing ZmPIN1a-YFP; arrowhead points to PIN1-YFP up-regulation at the site of primordia initiation. (Scale bars, 100 μm.)