Table 1.

Strains of Aspergillus niger used in this study

Strain of A. niger	Abbreviation	Description
AB1.13 pyrG+	AB1.13	Uridine prototroph of AB1.13 pyrG- [32]
AB1.13 CAD 4.1	AB1.13 CAD	Selected pyrG + transformant of cadA expressing transformant (CAD10.1) of AB1.13 [28]
AB1.13 CAD + MFS + MTT + CitB #99	CitB#99	citB overexpressing strain of AB1.13 CAD + MFS + MTT #49B [17]
AB1.13 CAD + MFS + MTT + CitB #99-acl A4 R AB1.13 CAD + MFS + MTT + CitB #99-acl C3 R AB1.13 CAD + MFS + MTT + CitB #99-acl D9 R AB1.13 CAD + MFS + MTT + CitB #99-acl E11 Z AB1.13 CAD + MFS + MTT + CitB #99-acl F11 Z AB1.13 CAD + MFS + MTT + CitB #99-acl G1 Z AB1.13 CAD + MFS + MTT + CitB #99-acl G8 Z AB1.13 CAD + MFS + MTT + CitB #99-acl G11 Z AB1.13 CAD + MFS + MTT + CitB #99-acl H4 Z AB1.13 CAD + MFS + MTT + CitB #99-acl A9 G AB1.13 CAD + MFS + MTT + CitB #99-acl A12 G AB1.13 CAD + MFS + MTT + CitB #99-acl B11 G AB1.13 CAD + MFS + MTT + CitB #99-acl D10 G AB1.13 CAD + MFS + MTT + CitB #99-acl D12 G	ACL A4 R ACL C3 R ACL D9 R ACL E11 Z ACL F11 Z ACL G1 Z ACL G8 Z ACL G11 Z ACL H4 Z ACL A9 Z ACL A12 G ACL B11 G ACL D10 G ACL D12 G	acl1 and acl2 overexpressing strain of AB1.13 CAD + MFS + MTT + CitB #99 (this study)