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. 2019 Oct 2;9:14212. doi: 10.1038/s41598-019-50572-8

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© The Author(s) 2019

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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The ACF-remodeled nucleosomes are associated with ~150 bp of DNA. (a) Enlarged view of an AFM topography image of a centrally positioned nucleosome and schematics of the analysis. (b) Two-dimensional histogram L_C/ΔL representing the complexed length of DNA L_C versus the nucleosome position ΔL. Left panel: nucleosomes incubated for 45 minutes at 30 °C in the absence of ACF (N = 1287 particles analyzed). Right panel: nucleosomes incubated with ACF (N = 4739 particles analyzed).