(A–F) Combined immunohistochemistry for β-III tubulin (Tuj1, brown) and in situ hybridization (blue) for the indicated probes (top panel and left four columns), or immunofluorescence for Prox1 or CK8, combined with immunofluorescence for β-III tubulin and a counterstain with DAPI (two right columns) in wild type (A,B,E) and double Neurog1/2 KO (C,D,F) at E14.5 or E16.5 as indicated. In the circumvallate papilla, markers are expressed on the dorsal surface (white arrowhead for CK8) but not at the lower part of the trenches (red arrowhead in the right column), where taste buds will develop after birth. CK8 is also expressed in flattened cells of the periderm (asterisks). For every probe two animals were examined. CP : circumvallate papilla; FP : fungiform papilla ; PP : palatal papilla ; R : ruga. Scale bars: 20 μm.