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. 2019 Oct 10;9:14540. doi: 10.1038/s41598-019-51101-3

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© The Author(s) 2019

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Bacterial twitching velocity with different pili distributions: (a) Twitching velocity for different pili distribution angles and pili numbers. As expected, the stationary time (V_t < 0.01 µm/s) decreases and moving time (V_t > 0.8 µm/s) increases as the number of pili increases because the bacterium is pulled by pili more frequently then. Pili distribution angle has significant influence on the intermediate velocity (0.01 < V_t < 0.8 µm/s); (b-c) The average twi_tching velocity is more sensitive to the number of pili than the pili distribution angle.