Evolutionary timescales of mandrill-speciation simian foamy viruses. The timescales were estimated under the FV–host co-speciation assumption by using the TDRP models: ; where and are time, and the corresponding number of substitutions per site, respectively. For envLP/5′SU, = 4.487 (95% HPD = 3.964–5.114), = 1.746 (95% HPD = 1.377–2.217); for envcenSU, = 4.114 (95% HPD = 3.827–4.383), = 1.762 (95% HPD = 1.556–1.985); for env3′SU/TM, = 4.522 (95% HPD = 4.098–4.973), = 1.553 (95% HPD = 1.273–1.892); and for pol, = 4.681 (95% HPD = 4.100–5.320), = 1.419 (95% HPD = 1.093–1.820). See Section 5 for the nodes used in the TDRP model calibrations.