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. 2019 Oct 22;9:15098. doi: 10.1038/s41598-019-51360-0

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© The Author(s) 2019

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Vegetative and flower morphology of the characters evaluated: (A) creeping habit in Anathallis; (B) caespitose habit with longer inflorescences than leaf in Pseudolepanthes; (C) proliferating ramicauls in Tubella; (D) ornamented lepanthiform bracts in Trichosalpinx; (E) laminar, motile lip (I) of Trichosalpinx; (F) bilobed stigma and glenion (G) in Lepanthopsis; (G) Appendix (A) at the lip base of Lepanthes; (H) Column foot (cf) and ventral anther in Gravendeelia; (I) Bilobed lip (B) and apical anther in Lepanthes; (J) Ventral anther (an) and stigma (S) in Anathallis; (K) Pollinarium with viscidium (V) and caudicles (C) in Lepanthe; (L) Pollinarium with caudicles (C) in Trichosalpinx. Photographs (A–I) by D.Bogarín, (B) by S. Vieira-Uribe.