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NMR backbone amide CSP profiles of CXCL1 on adding heparin (A), HS (B), CS (C), and DS (D). Basic and buried residues that show significant perturbation are shown in blue and red, respectively. The chemical shift of Lys-21 (green asterisks) is perturbed by 1.20, 0.95, 0.26, and 0.75 ppm for heparin, HS, CS, and DS, respectively.
