Energy-level diagram. The localization of the S1 and S2 energy levels of chlorophyll a (Chl a) and S1 energy levels of carotenoids. The energies of Chl a: S2(Qx) 16 000/cm and S1(Qy) 14 700/cm. The energies of the S1 state of xanthophylls: violaxanthin (Vx, 15 290/cm), diadinoxanthin (Ddx, 15 130/cm), antheraxanthin and lutein (Ax/L, 14 720/cm), diatoxanthin (Dtx, 14 485/cm), and zeaxanthin (Zx, 14 170/cm).28 Arrows from left to right represent forward energy transfer (light-harvesting); arrows from right to left – reverse energy transfer (NPQ).