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. 2019 Nov 7;10:5068. doi: 10.1038/s41467-019-12927-7

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Fig. 7 — Model for Ug99 origin by somatic hybridisation and nuclear exchange. The ancestral isolate of the Ug99 lineage acquired the A and C genomes from an isolate of the 21 lineage and an unknown isolate and later gained virulence to wheat cultivars carrying the Sr31 resistance gene