Table I.

Symbol	Physiological parameters	Dimension	Value
Geometrical
(Mason and Larkman,1990; Kawaguchi,1993; Mainen et al.,1995; Larkman and Mason,1990; Wong et al.,2000)
A IN	Area of Soma (GABAergic IN)	cm 2	∼0.908 × 10−5
A PC	Area of Soma (layer V PC)	cm 2	1.393 × 10−5
A ${}_{\text{PC}}^{\text{B}}$	Area of Basal (layer V PC)	cm 2	∼18 × 10−5
A ${}_{\text{PC}}^{\text{A}}$	Area of Apical Tuft (layer V PC)	cm 2	∼6 × 10−5
l BS	Basal Stem Length “Maximal Number of Intersections”	cm	0.6 × 10−2
l AS	Apical Stem Length	cm	∼6 × 10−2
S PC	Mean Dendrite Transversal Area	cm 2	∼0.44 × 10−8
Electrophysiological
(Major et al.,1994; Spruston et al.,1994; Kleber and Riegger,1987)
c m	Membrane Capacitance	μF/cm 2	0.75
r m	Membrane Resistance	Ω cm 2	40000
τm = c m r m	Membrane Time Constant	ms	30
r i	Intra‐cellular Axial Resistance	Ω cm	166
αPC	Layer V PC Synaptic Factor	pA (0 ∼ 1.5)	0.4
αIN	GABAergic IN Synaptic Factor	pA (0 ∼ 1.5)	0.3
r e	Extra‐cellular Resistance	Ω cm	63
Vascular
(Friston et al.,2000)
χPC	Layer V PC Energetic Factor	nM (0 ∼ 1)	1.0
χIN	GABAergic IN Energetic Factor	nM (0 ∼ 1)	0.8
τs	Signal Decay	ms	1540
τf	Autoregulation	ms	2460
τ0	Transit Time	ms	980
α	Stiffness	Dimensionless	0.33
E 0	Resting Oxygen Extraction Fraction	Dimensionless	0.34