Table 1. Main model parameters and their values unless otherwise noted.

Values that change between runs, idle periods, and allocentric corrections are indicated accordingly.

Parameter	Variable	Value
Neurons per population	$n\times n$	$232\times 232$
Simulation timestep	$\mathrm{\Delta }t$	1 ms
Exc. membrane time constant	${\tau }_{\text{m}}^{+}$	40 ms
Inh. membrane time constant	${\tau }_{\text{m}}^{-}$	20 ms
Exc.-to-exc. synaptic delay	${\tau }_{\text{s}}^{++}$	5 ms
Exc.-to-inh. synaptic delay	${\tau }_{\text{s}}^{-+}$	2 ms
Inh. synaptic delay	${\tau }_{\text{s}}^{-}$	2 ms
Exc. drive maximum	$a_{\text{max}}^{+}$	$\{\begin{array}{cc}2.0\hfill & \text{runs}\hfill \\ 1.6\hfill & \text{idle}\hfill \\ 2.0\hfill & \text{allo.}\hfill \end{array}$
Exc. drive minimum	$a_{\text{min}}^{+}$	0.8
Exc. drive scaled spread	${\rho }_{a^{+}}$	$\{\begin{array}{cc}1.2\hfill & \text{runs}\hfill \\ 0.9\hfill & \text{idle}\hfill \end{array}$
Inh. drive magnitude	$a_{\text{mag}}^{-}$	$\{\begin{array}{cc}0.72\hfill & \text{runs}\hfill \\ 0.0\hfill & \text{idle}\hfill \\ 0.72\hfill & \text{allo.}\hfill \end{array}$
Inh. drive theta amplitude	$a_{\text{th}}^{-}$	$\{\begin{array}{cc}0.2\hfill & \text{runs}\hfill \\ 0\hfill & \text{idle}\hfill \\ 0\hfill & \text{allo.}\hfill \end{array}$
Inh. drive theta frequency	$f$	8 Hz
Exc. synaptic strength	$w_{\text{mag}}^{+}$	0.2
Exc. synaptic spread	$r_{w^{+}}$	6
Inh. synaptic strength	$w_{\text{mag}}^{-}$	2.8
Inh. synaptic distance	$r_{w^{-}}$	12
Exc. synaptic shift	$\xi$	3
Exc. velocity gain	$\alpha$	0.25 s/m
Exc. noise magnitude	${\displaystyle \mathrm{V}\mathrm{a}\mathrm{r}[{\zeta }^{\mathrm{P}}(\mathbf{r},\mathrm{t})]}$	0.0022
Inh. noise magnitude	${\displaystyle \mathrm{V}\mathrm{a}\mathrm{r}[{\zeta }^{-}(\mathbf{r},\mathrm{t})]}$	0.0022