Fig. 4. Combined environmental and genetic effects account for variation in the start of torpor.

Plots show correlation between fitted and observed values for the start of torpor (in days from hibernaculum placement) using a linear regression model with n = 72 independent squirrels. Adjusted R²-value is labeled in each. Shading matches Fig. 2a, c. a Linear model fit with the environmental variables of year of monitoring and date of hibernaculum placement, and the biological variable of sex (i.e. the fixed effects, see methods). b Linear model fit with the 14 significant and suggestive SNP genotype combinations for each squirrel. c Linear model fit with the variables from both a and b.