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. 2019 Dec 27;9:20089. doi: 10.1038/s41598-019-56577-7

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© The Author(s) 2019

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Zinc distribution in midgut gland sections of zinc-exposed Arion vulgaris with colour dithizone staining (a,b) and fluorescent toluenesulfonamidoquinoline (TSQ) staining (c). Midgut gland cross sections showing lumen (LU) surrounded by digestive cells (DC) and calcium cells (CC) containing calcium granules (GR). Zinc stained by dithizone (red colour) (a,b) is exclusively allocated in calcium cells. Fluorescent staining by TSQ localize zinc (bright blue colour) (c) mainly in cytoplasm of calcium cells and on the outer edge of calcium granules (c - GR). The bar corresponds to a size of 50 µm.