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. 2019 Dec 27;9:20152. doi: 10.1038/s41598-019-56817-w

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© The Author(s) 2019

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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LKU4 administration induces browning of iWAT in mice fed a HFD. (A,B) The expression of browning genes in iWAT was determined by RT-qPCR (A) and immunoblotting (Cropped blots were used). (B–D) Mitochondrial DNA (mtDNA) copy number and citrate synthase (CS) activity (C) and representative images of VDAC2 and UCP1 staining (D) in iWAT of HFD and HFD-LKU4 mice. (E) Rectal temperatures were measured in each group of mice at week 12 (n = 8–10). HFD vs HFD-LKU4; *P < 0.05, **P < 0.005.