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Benzyl glucuronoate trapped on the surface of the S267A OtCE15A variant (PDB code 6T0E). Presumably, the cleavage rate was too fast to capture the Michaelis–Menten complex with benzyl glucuronoate over the short crystal soaking period (10 s) but allowed capture of the uncleaved substrate along the edge of the active site cleft ∼10 Å from the catalytic center (glucuronate portion in green and benzyl portion in magenta).
