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. 2020 Jan 31;10:1546. doi: 10.1038/s41598-020-58150-z

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© The Author(s) 2020

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Binding-analysis of a CD40/OX40 specific tetra-VH IgG. The antibody was evaluated for (a) binding to recombinant proteins in ELISA and for binding to (b) transfected cells or (c) endogenously expressing cells in flow cytometry. In vitro activated (IVA) CD4+ T-cells express OX40 and 4-1BB and the B-cell line Raji express CD40.