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. 2020 Feb 6;10:1991. doi: 10.1038/s41598-020-58522-5

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© The Author(s) 2020

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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A1S_3296 promoted biofilm formation through Csu pili mediated pathway. The effect of A1S_3296 in csuA and pilA mutants regarding congo red binding in macro colonies (A) and biofilm formation. (B) Immunoblot blot to illustrate alteration in CsuAB expression in 2396 mutant as compare to wild type and complemented strain. (C) The strain lacking the expression of csuABC operon (∆csuA) was used as negative control for CsuAB expression.