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. 2020 Feb 13;10:2537. doi: 10.1038/s41598-020-59488-0

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© The Author(s) 2020

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Phylogeny of Rickettsia constructed using maximum-likelihood (ML) estimations based on concatenated 16S rDNA, gltA and atpA sequences (1886 unambiguously aligned nucleotide sites; best-fit approximation for the evolutionary model: GTR + G + I). Sequences from Rickettsia characterized in this study are shown in red. Only one 16S rDNA, gltA and atpA concatenated sequence per Rickettsia genotype and per tick species is shown. Sequences from representative Rickettsia groups, species and strains available in GenBank were also added to the analysis. The grey boxes delineate the different Rickettsia groups (their names are indicated in upper case), including the novel guiana group described in this study. Bacterial name, host species and GenBank accession numbers are shown on the tree. Branch numbers indicate percentage bootstrap support for major branches (1000 replicates; only bootstrap values >70% are shown). The scale bar is in units of substitution/site.