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. 2020 Feb 13;8:e8519. doi: 10.7717/peerj.8519

Table 1. Additions and adaptations to the original mesowear technique-ordered by mesowear technique and date.

Technique References Description Scores
Original mesowear—Mesowear I Fortelius & Solounias (2000) –Using the naked eye or ×10 magnification OR: low, high
–Scoring only sharpest buccal cusp of maxillary M2 CS: blunt, round, sharp
–Last molar in occlusion and M1 shape similar to M2
–Percentage of high relief and Percentage of sharp, round and blunt cusps
Mesowear I—Adapted for Equidae Kaiser & Fortelius (2003) Method extended to all apices on maxillary P4–M3 in equids Replaces original mesowear
Mesowear I Franz-Odendaal & Kaiser (2003) Method extended to maxillary M3, and mandibular M2 in ruminants Replaces original mesowear
Mesowear I—Adapted for Lagomorpha Fraser & Theodor (2010) ‘Cusp relief’ combined with ‘buccal shearing crush wear’ on maxillary and mandibular P4–M2–resulting in 5 dietary classes 1: 45° enamel-dentine relief with no additional wea
2: 45° enamel-dentine relief with buccal shearing crush wear
3: 45° enamel-dentine relief with buccal shearing crush & phase II wear
4: 90° enamel-dentine relief with no additional wear
5: 90° enamel-dentine relief with buccal shearing crush wear
Mesowear I—Adapted for Conodonta Purnell & Jones (2012) Scored on P1 elements Not truly mesowear, does not have scores
Mesowear I—Adapted for Leporines & Murines Ulbricht, Maul & Schulz (2015) Classical mesowear on the maxillary M1–M2, and mandibular p3 in leporinae and distal side of the maxillary M1 and mandibular m1 in murinae Same scores as original mesowear
Mesowear I—Adapted for voles Kropacheva et al. (2017) Maxillary M1–M2, mandibular m1 Occlusal relief 1–7
Lateral facet development 1–3
Mesowear II—‘Mesowear ruler’ Mihlbachler et al. (2011) Simplified score using gauges and a seven-point system Combined score 0–6
Mesowear II—‘Mesowear ruler’ Wolf, Semprebon & Bernor (2012) Additional intermediate scores Combined score 0–13 in increments of 0.5
‘Mesowear angles’—Adapted for Proboscidea Saarinen et al. (2015) Mean mesowear angles of three central lamellae in occlusion’ on all except deciduous teeth Mean mesowear angle
<106°: C3-plant based diet
>130°: C4-plant based diet (grazer)
“Mesowear angles”–Adapted for Xenarthra Saarinen & Karme (2017) All molariform teeth For Xenarthra, Folivora:
Mean mesowear angle:
60°–85°: fruit browsers
75°–100°: leaf browsers
100–132°: mixed-feeders
132°–150°: grass dominated mixed-feeders
150°–190°: grazers
For Xenarthra, Cingulata:
60°–100°: carnivore, insectivore, omnivore, possibly browsers
100°–125°: browse-dominated mixed-feeders & herbivorous omnivores
125°–152°: grass-dominated mixed-feeder
152°–190°: grazers
Mesowear II Mihlbachler & Solounias (2006) Simplified score, only proportion of sharp cusps Proportion of sharp cusps:
40–100%: Clean browser
20–40%: Mixed feeders:
0–20%: Grazer
Mesowear II ‘quantitative mesowear’ Widga (2006) Interval measurements of cusp and saddle heights to calculate cusp relief Index of cusp relief:
Low ICR: grazer
High ICR: browser
Mesowear II Rivals & Semprebon (2006) Simplified score combining OR and CS 0: high relief & sharp cusps
1: high relief & round cusps
2: low relief & round cusps
3: low relief & blunt cuspss
Mesowear II Kaiser (2009) 0: high relief & sharp cusps
1: high relief & round cusps
2: low relief & sharp cusps
3: low relief & round cusps
4: low relief & blunt cusps
Mesowear II Rivals, Schulz & Kaiser (2009) 0: high relief & sharp cusps
1: high relief & round cusps
2: low relief & round cusps
2.5: low relief & sharp cusps
3: low relief & blunt cusps
Mesowear II Croft & Weinstein (2008) 0: high relief & sharp cusps
1: high relief & round cusps
2: low relief & round cusps
2.5: low relief & sharp cusps
3: high/low relief & blunt cusps
Mesowear II Fraser et al. (2014) Method extended to mandibular P4–M3 for ruminants 1: high relief & sharp cusps
2: high relief & round cusps
3: high relief & very round cusps
4: low relief & round-blunt cusps
5: low relief & flat-blunt cusps
Mesowear II—Adapted for Marsupialia Butler, Louys & Travouillon (2014) Use of classical mesowear and a combined score on the maxillary left maxillary molars, scoring sharpest buccal cusp Combined score as in Kaiser (2009)
Mesowear I & II—Expanded Winkler & Kaiser (2011) Intermediate stages added to original and combined score OR: low, high-low, high, high-high
CS: blunt, round-round, round, round-sharp, sharp
Combined score 1–17
Mesowear I and II—Expanded, Adapted for Rhinocerotidae Taylor et al. (2013) Expanded version and combined score on maxillary P2–M2. Combined score 1–11
Mesowear III—‘Inner mesowear’ Solounias et al. (2014) Scores the second enamel band, using a stereo-microscope
Mesial side, distal side and junction point are scored separately
Enamel band wear states:
1: ideal browser
2–3: intermediate
4: ideal grazer
Junction point score 1–4