Table 1.

Summary of definitions of terms used in the four-locus population genetic model.

T2 / T1 ; t1 / t2	Allele for attractive display trait / allele for unattractive display trait, expressed by males; lowercase denotes the respective allele frequencies
C2 / C1 ; c2 / c1	Allele for the capacity to attempt forced fertilization (coercion) / Allele for the lack of capacity to coerce, expressed by males; the respective allele frequencies
P2 / P1 ; p2 / p1	Allele for a biased behavioral mating preference for males with the T2 allele by a factor (1+a) / Allele for unbiased mating, expressed by females; the respective allele frequencies
R2 / R1 ; r2 / r1	Allele for the capacity to resist attempted forced fertilization / Allele for the lack of capacity to resist attempted forced fertilization, expressed by females; the respective allele frequencies
st	Viability cost to males with the T2 allele of producing an attractive display trait
sc	Viability cost to males with the C2 allele associated with the capacity to coerce females
1+a	Factor by which females with the P2 allele prefer to mate with males having the T2 allele
b	Likelihood of a male with the C2 allele to succeed in forced fertilization of a female that lacks resistance (has the R1 allele)
γ	Effectiveness of the capacity to resist attempted forced fertilization in females having the R2 allele.