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Transmembrane mucins emanating up to 500 nm above the plasma membrane interact with multimeric galectin-3 to promote epithelial integrity. Major classes of glycans at the ocular surface include O-glycans, commonly attached to serine (Ser) or threonine (Thr) amino acids, and N-glycans, attached to asparagine (Asn). Mucins, glycoproteins and glycolipids are embedded in the plasma membrane.
