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. 2020 Feb 14;2020:3535982. doi: 10.1155/2020/3535982

Table 2.

Effect of probiotic supplementation in CRC experimental animals.

Experimental model Intervention Duration of treatment Key results References
ApcMin/+ CRC mouse model L. acidophilus ATCC 314, L. fermentum NCIMB 5221 (each 0.5 × 1010 CFU; total 1 × 1010 CFU/day) 12 weeks ↓ multiplicity of tumors
β-catenin and Ki-67
[29]

Azoxymethane-mediated colonic neoplasm induced Sprague-Dawley rats B. lactis (1 × 1011 CFU/g), and/or resistant starch (Hi-maize® 958 or Hi-maize® 260; 100 g/kg diet) ∼22 weeks ↓ incidence and development of colonic neoplasms
The protective effects were observed to be higher in the synbiotic supplemented group
[33]

Trinitrobenzene sulfonic acid-mediated chronic colitis induced Sprague-Dawley rats VSL#3 (B. breve, B. infantis, B. longum, L. acidophilus, L. bulgaricus, L. casei, L. plantarum, and Streptococcus salivarius subsp. thermophilus), 5 × 109 CFU/100 g of body weight Differs∗∗ No carcinoma development
No high-grade dysplasia
↓ colon damage
↑ expression of antiangiogenic factor angiostatin, alkaline sphingomyelinase, and vitamin D receptor.
[34]

Azoxymethane/dextran sodium sulfate-mediated colitis-associated CRC induced mouse model VSL#3 (1.3 × 106 CFU/day), and/or Balsalazide (300 mg/kg body weight/day) 2 weeks before azoxymethane exposure and continued for 9 weeks until sacrification ↓ number of tumors
↓ F4/80-positive macrophages
↓ p-STAT3 expression
↓ BCL-2 expression
↓ MIP-1β, MCP-1, IL-6, IL-10 level
↑ BAX expression
[35]

Azoxymethane-mediated colitis-associated CRC induced mouse model VSL#3 (1 × 109 CFU/day) 19 weeks (from 6th week to 24th week) Clostridium species
No reduction in tumorigenesis
[36]

1,2-Dimethyl hydrazine (DMH)-mediated CRC induced Sprague Dawley rats L. rhamnosus GG, or L. casei, L. plantarum, or L. acidophilus, or B. bifidum. (probiotic dose: 1 × 109 CFU/day) Seven weeks (1 week before starting DMH exposure and continued for 6 weeks) ↓ percentage of Aberrant crypt foci (ACF)
↓ nitroreductase activity, β-glucuronidase activity, β-glucosidase activity
[37]

DMH-mediated CRC-induced Sprague Dawley rats Synbiotic (L. rhamnosus GG, L. acidophilus, and inulin; 1 × 109 CFU probiotic +5 mg inulin/day) or probiotic (L. rhamnosus GG, and/or L. acidophilus; 1 × 109 CFU probiotic/day) or prebiotic (inulin; 5 mg/day) 19 weeks (1 week before starting DMH exposure and continued for 18 weeks) ↓ MDA level
↑ GSH, SOD, and GPx
Improved the histopathological score
[38]

DMH-mediated CRC-induced rats L. acidophilus LaVK2 and B. bifidum BbVK3 or both probiotic + piroxicam; 2 × 109 CFU/g of each probiotic 32 weeks ↓ number of ACF, mucin-depleted foci, and proliferating cell nuclear antigen [39]

DMH-mediated CRC-induced rats L. plantarum AS1 (1 × 109 CFU/day) 5–21 weeks ↑ antioxidant system of the host
↓ tumor diameter and number of tumors
[40]

DMH-mediated CRC-induced rats L. salivarius (5 × 108 or 1 × 1010 CFU/kg body weight/day) 15 weeks (2 week before starting DMH exposure and continued until 15 weeks) Improved the colonic microflora and luminal metabolisms.
↓ number and multiplicity of ACF, azoreductase activity
↑ short-chain fatty acid levels
[41]

CT26 tumor-bearing mice L. plantarum or L. rhamnosus; 1 × 109 CFU/day Pre-exposure for 14 days ↓ CT26 growth
↑ lifespan of tumor-bearing mice
↑ IFN-γ, Th1-type CD4+ T differentiation
↑ CD8+ function
↑ NK cell infiltration
[42]

DMH-mediated CRC-induced rats L. salivarius Ren (5 × 1010 CFU/kg body weight/day) 32 weeks Reversed the DMH-induced altered microbiota [43]
DMH-mediated CRC-induced rats L. rhamnosus GG CGMCC 1.2134 (1 × 109 CFU/day) 25 weeks ↓ incidence, multiplicity, and volume of tumor
↓expression of inflammatory proteins, and antiapoptotic protein
↑ proapoptotic proteins
[44]

DMH-mediated CRC-induced mice L. casei BL23 (10 µl; 1 × 108 CFU/µl) 3 days (on days 0, 14, 28) ↓ incidence of tumor
↓ multiple plaque lesions
Regulates the Treg and Th17 T cells
Altered the expression of IL-6, IL-10, IL-17, and TGF-β
[45]

DMH and SW480 cell-mediated CRC-induced rat B. infantis (1 × 109 CFU/day) and/or 5-FU + oxaliplatin 11 days ↑ body weight and intestinal villus height
↓ IL-6, IL-1β, TNF-α levels, and Th17 and Th1 cell-associated cytokines
↑ CD4+, CD25+, Foxp3+, Tregs expressions
[46]

Azoxymethane/dextran sodium sulfate-mediated colitis-associated cancer-induced mice model B. longum, L. acidophilus, and E. faecalis (1.2 × 107 CFU/day) Pretreatment for 2 weeks and continued till the end of the experiment ↓ intestinal inflammation and tumor formation.
Desulfovibrio, Mucispirillum, and Odoribacter species
Lactobacillus species
Altered the expression of CXCR2 ligand genes
[47]

Azoxymethane-mediated CRC-induced mice L. acidophilus (1 × 109 CFU/day) and B. bifidum (1 × 109 CFU/day) 5 months ↓ miR-135b, miR-155, and KRAS ↑ miR-26b, miR-18a, APC, PU.1, and PTEN [48]

Azoxymethane/dextran sodium sulfate-mediated CRC-induced rat model L. acidophilus LA5 and/or B. animalis subsp. Lactis BB-12, and GBR; 5 × 107 CFU of single probiotic strain/day or 2.5 × 107 CFU each strain/day 10 weeks ↓ mucin-depleted foci formation
↑ expression of p53, Bax, caspase-3, and Bax/Bcl-2 ratio
↓ Bcl-2 expression
↑ SOD activity
↓ aberrant crypt foci (ACF)-producing sialomucin
[49]

DMH-mediated CRC-induced rats L. rhamnosus GG MTCC #1408, and/or L. acidophilus NCDC #15 (1 × 109 CFU/day), and/or celecoxib (6 mg/kg body weight) 18 weeks ↓ multiplicity and tumor burden
↓ Bcl-2, K-ras expression
↑ Bax, p53 expression
[50]

DMH-mediated CRC-induced rats L. rhamnosus GG MTCC #1408, and/or L. acidophilus NCDC #15 (1 × 109 CFU/day), and/or celecoxib (6 mg/kg body weight) 6 weeks ↓ ACF formation
β-catenin, COX-2, and NF-κB expression
[51]

↑: increased; ↓: decreased; MDA: malondialdehyde; GSH: glutathione; SOD: superoxide dismutase; GPx: glutathione peroxidase; NK: natural killer; IFN-γ: interferon-γ; p-STAT3: phospho-signal transducer and activator of transcription 3; BCL-2: B-cell lymphoma 2; BAX: BCL2-associated X protein; MIP-1β: macrophage inflammatory protein 1 beta; MCP-1: monocyte chemoattractant protein-1; IL-6: interleukin-6; IL-10: interleukin-10; KRAS: Kirsten rat sarcoma 2 viral oncogene homolog; GBR: germinated brown rice; Hi-maize® (high-amylose maize starch was used as a source of resistant starch); ∗∗From one week before colitis induction to death of the experimental animal.