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. 2020 Mar 2;15(3):e0229204. doi: 10.1371/journal.pone.0229204

Table 2. Alpha diversity measures and descriptions.

Alpha Diversity Measure Description Citations
Ace An estimate of species richness using a correction factor [18,34]
Observed OTUs The number of different OTUs per sample [16]
Chao1 A measure of species richness, which gives more weight to rare species [34]
Margalef Measure of species richness in a given area or community [35,36]
Fisher alpha Relationship between the number of species and the relative abundance of each species; referred to as a measure of richness [37,38]
Faith pd Sum of OTU branch lengths; takes into account phylogenic distance between OTUs; the greater the number of unique OTUs, the higher this index will be [18,39]
Brillouin d Measures the diversity of the species present [18]
Shannon Calculation of richness and evenness using a natural logarithm; accounts for both relative abundance and evenness of the taxa present; referred to as an attempt for one metric to capture both richness and relative evenness [18,40]
Enspie (ENS) Measure of the number of equally abundant species; if the relative abundance of all species in a community were exactly identical, ENS would simply be the total number of species in that community [41]
Menhinick The ratio of the number of taxa to the square root of the sample size; referenced as "species richness" [18,35]
Mcintosh e A measure of how evenly/unevenly abundant taxa are in the sample [18,42]
Simpson Measure of relative abundance of the different species making up the sample richness; referred to as an attempt for one metric to capture both richness and relative abundance [40,43]
Berger Parker d A measure of relative richness of the abundant species; higher values indicate relative abundance disturbance [18,44,45]
Strong Assesses species relative abundance unevenness/dominance concentration [46]
Simpson e A measure of evenness based on the number of organisms and number of species [43]
Pielou e Measure of relative evenness of species richness [12,42,47]
Heip e A measure of evenness [18,42]
Lladser pe A measure of how much of the environment contains unsampled taxa [18,48]