TABLE 2.

Permutational multivariate analysis of variance (PERMANOVA) with nested design according to the horizontal and vertical positions of Cladonia squamosa.

Ecological factors	Df	Algae		Lichen-associated fungi		Bacteria
		F ratio	P-value	F ratio	P-value	F ratio	P-value
Horizontal position	2	0.245	0.8854	0.435	0.8977	0.482	0.741
Vertical position nested in horizontal position	2	2.524	0.0197	2.759	0.0001	4.958	0.0001