Table 1.

List of notations.

Notations	Meaning
θ	Spike event
x	The sum of pre-spike events over time
w	Synaptic weight
Vmem	Membrane potential
Vth	Neuronal firing threshold
I	Input current at each time step
net	Total current influx over time
a	Activation of spiking neuron
E	Loss function
δ	Error gradient