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. 2020 Mar 12;10:4556. doi: 10.1038/s41598-020-61279-6

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© The Author(s) 2020

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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A crateriform PPs in the midpiece colon of a 42-day old Duan-Nai-An-treated piglet (H&E staining). The inner structure of the PPs appeared scrotiform with well-developed lymphoid tissues on the lateral and basal surfaces of the inner wall of the cyst, well-developed lymphoid follicles near the circular muscle layer, and a large number of scattered lymphocytes in the lamina propria. LP indicates Lamina propria; MM indicates muscularis mucosae; MW indicates muscular wall; PPs indicate Peyer’s Patches; and S indicates Submucosa.