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. 2020 Mar 13;10:322. doi: 10.3389/fonc.2020.00322

Table 4.

Therapeutic compounds used for autophagy induction or inhibition in cancer.

Compound Target Model system Autophagy evaluation References
GSK2606414 PERK Basal cell carciona (BCC/KMC1)
Gastric Adenocarcinoma (AGS)
Melanoma (A375)
Imiquimod
↓LC3-II
↓EGFP-LC3-II puncta
(71)
Celastrol mTOR Glioma (U251, U87 and C6)
Pre-treatment CQ
↑LC3, BECN1,p62
↑LC3 puncta
≠LC3-II
(85)
LY294002
UCN-01
(7-hydroxystaurosporine)
PI3K
Akt
Glioma (U87-MG, U373-MG and T98G)
Rapamycin
MDC stain (86)
Rapamycin
RAD001
(rapalogue)
KU-0063794
(catalytic mTOR inhibitor)
WYE-354
(catalytic mTOR inhibitor)
mTOR Bladder carcinona (RT112)
osteosarcoma (U2OS)
neuroblastoma (SK-N-SH)
squamous cell carcinoma (HN10)
↑LC3 punctate (all treatments)
↑LC3 punctate (only with catalytic mTOR inhibitors)
(87)
AZD8055 mTOR lung cancer
H838 and A549
+E64d/pepstatinA
↑LC3-II ↑LC3 puncta
↑Acridine orange stain
↑↑LC3-II
(93)
Metformin AMPK Melanoma (A375, and SKMel28)
Xenograft model
↑LC3-II, BECN1 ↑LC3 puncta ↑Autophagosomes (TEM)
↑ CatB activity
↑LC3-II ↑LC3 puncta
(94)
Metformin mTOR Esophageal squamous cancer cells (ESCC)
Pre-treatment 3-MA or CQ
Xenograft
↑LC3-II, BECN1
↑GFP-LC3 puncta
↑ Autophagosomes (TEM)
↑ Acridine orange and MDC stain
↓LC3-II, BECN1
↓LC3-II, ↓p62 (IHC)
(95)
Akt inhibitor
(1L-6-hydroxymethyl-chiro-inositol 2(R)-2-O-methyl-3-O-octadecylcarbonate)
AKT Glioma
U87-MG
U87-MG ΔEGFR
↑GFP-LC3 punctate ↑autophagosomes (TEM), ↑Acridine orange stain (91)
Curcumin Akt/mTOR/p70S6K/4E-BP Glioma
U87-MG and U373-MG
Xenograft
↑GFP-LC3 puncta ↑LC3-II ↑autophagosomes (TEM) ↑ Acridine orange stain
↑LC3-II, ↑LC3 (IHC)
(92)
Quercetin Akt-mTOR Breast cancer
MCF-7 and MB-231
↑LC3-II, LC3 puncta (89)
Quercetin Akt-mTOR Gastric adenocarcinoma AGS and MKN28 ↑ LC3-II, BECN1, ATG7, ATG5/12 ↑ GFP-LC3 puncta
↑ Autophagosomes (TEM) ↑ Acridine orange stain
(88)
AZD1208 pan-PIM kinase inhibitor Chronic lymphocytic leukemia (CLL)
BafA1
↑Acridine orange stain, ↑LC3-II
↑↑LC3-II
(96)
Parthenolide Oxidative stress downregulation of 4E-BP1 See Table 2 See Table 2 (38)
Selenite eIF2alpha phosphorylation by ROS or ER stress See Table 2 See Table 2 (41, 42)
Erastin Ferroptosis inducer See Table 2 See Table 2 (49, 54)