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. 2020 Mar 24;10:5334. doi: 10.1038/s41598-020-62073-0

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© The Author(s) 2020

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Model of biennial mast cycle in European beech. Basic biennial mast cycle of European beech with weather-driven disturbances in spring leading to pollination failure (dashed black arrow) and, newly postulated, in summer due to extremely hot and dry conditions leading to fruit abortion (red arrow). ‘On’ year = potential mast year, ‘Off’ year = year after successful mast year, without flower buds. Scheme according to Lavee (2007)⁴⁰. Original artwork by Anita Nussbaumer.